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Biostratigraphy and systematics of late Asselian–early Sakmarian (Early Permian) fusulinids (Foraminifera) from southern Turkey

Published online by Cambridge University Press:  19 February 2008

CENGİZ OKUYUCU
CENGİZ OKUYUCU*
Affiliation:
Maden Tetkik ve Arama Genel Müdürlüğü (MTA), Jeoloji Etütleri Dairesi, 06520 Balgat, Ankara, Turkey
*
*E-mail: okuyucu@mta.gov.tr
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Abstract

The Anatolian Platform, which was a part of the Gondwanan Platform, is mainly characterized by carbonate-dominated deposits ranging in age from Devonian to Permian. The biostratigraphy and systematics of a late Asselian–early Sakmarian fusulinid fauna from the Anatolian Platform including Eastern and Central Taurides have been investigated in three sections: Özbek Hill, Eskibey and Bademli. Twenty-four fusulinid taxa, belonging to twelve genera, were determined in a single fusulinid zone dated as late Asselian–early Sakmarian. Early–middle Asselian fusulinid faunas have not been observed in any of the measured sections throughout the Anatolian Platform. This indicates that lower to middle Asselian deposits are represented by an interval characterized by quartz sandstone overlying upper Gzhelian strata. Five new species (Pseudochusenella anatoliana, Pseudofusulinoides altineri, Pseudofusulinoides convexus, Pseudofusulinoides subglobosus and Pseudofusulinoides vachardi) are described in this study. The Early Permian fusulinid fauna correlates very well with the fauna of other sections in the Palaeotethyan realm (Southern Alps, Central Asia, Southern China and Japan).

Keywords

biostratigraphyfusulinidsEarly PermianAnatolian PlatformTurkey
Type
Original Article
Information
Geological Magazine , Volume 145 , Issue 3 , May 2008 , pp. 413 - 434
Copyright
Copyright © Cambridge University Press 2008

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References

Bensh, F. R. 1962. Late Carboniferous and Lower Permian fusulinids of Northern Fergana. In Stratigraphy and Paleontology of Uzbekistan, Book 1, pp. 187–252. Akademiya Nauk Uzbekskoi SSR.Google Scholar
Bensh, F. R. 1972. Stratigraphy and fusulinids of Upper Palaeozoic of the southern Fergana (Central Asia). Akademiya Nauk Uzbekskoi SSR, Institut Geologii i Geofizikii, Izdatelstvo ‘Fan’ Tashkent, 1–147.Google Scholar
Bensh, F. R. 1987. Systematic revision of the pseudofusulinid genus Pseudofusulina Dunbar and Skinner and similar genera. Voprosy Mikropaleontologii 29, 20–53.Google Scholar
Blumenthal, M. M. 1944. Schichtfolge und -bau der Taurusketten im Hinterland von Bozkır. Revue de la Faculty des Sciences de la Université d'Istanbul, Serie B 9 (2), 95–125.Google Scholar
Blumenthal, M. M. 1951. Recherches géologieques dans le Taurus occidental dans l'arriére-pays d'Alanya. Mineral Research and Exploration Publications, Serie D 5, 134 pp.Google Scholar
Brunn, J. H., Dumont, J. F., Graciansky, P. C., Gutnic, M., Juteau, T., Marcoux, J., Monod, O. & Poisson, A. 1971. Outline of the geology of the Western Taurids. In Geology and History of Turkey (ed. Campbell, A. S.), pp. 225–55. Petroleum Exploration Society of Libya, Tripoli.Google Scholar
Brunn, J. H., Argyriadis, I., Marcoux, J., Monod, O., Poisson, A. & Ricou, L. 1973. Antalya'nin ofiyolit naplarinin orijini lehinde ve aleyhinde kanitlar. Cumhuriyetin 50. Yili Yerbilimleri Kongresi, MTA Publication, Ankara, 58–69.Google Scholar
Chang, L. H. 1963. Upper Carboniferous fusulinids from Kelpin and its neighbourhood, Hsin-Kiangs. Part I. Acta Palaeontologica Sinica 11 (1), 36–55.Google Scholar
Chen, S. 1934. Fusulinidae of South China, Part. 1. Palaeontologica Sinica, Series B 4 (2), 1–185.Google Scholar
Chen, X. & Wang, J. H. 1983. The fusulinids of the Maping Limestone of the Upper Carboniferous from Yishan, Guangxi. Palaeontologica Sinica, N. S. B. 19, 1–139.Google Scholar
Choi, D. R. 1973. Permian fusulinids from the Setamai–Yahagi district, southern Kitakami Mountains, Northeast Japan. Journal of the Faculty of Science, Hokkaido University, Series 4 16 (1), 1–132.Google Scholar
Davydov, V. I. 1980. Rugosofusulininae – a new fusulinid subfamily with a rogose wall. Materials of the VIII Micropaleontology Meeting, Baku, “Elm”, 44–5.Google Scholar
Davydov, V. I. 1986. About a phylogenetic criterion of weighing specific features in foraminifer systematics (exemplified by fusulinids). Benthos ‘86, Resumes Abstracts, Genéve, Museum d'Histoire Naturelle, 35.Google Scholar
Davydov, V. I., Popov, A. V., Bogoslovskaya, M. V., Chernykh, V. V., Kozitskaya, R. I. & Akhmetshina, L. Z. 1993. Aidaralash section. In Permian system: guide to geological excursion in the Uralian type localities (eds Chuvashov, B. I. & Nairn, A. E. M.), pp. 131–46. Occasional Publications, Earth Science and Research Institute, New Series 10.Google Scholar
Demirel, I. H. & Tekin, U. K. 1993. Gondvanya Platformu Paleozoyik-Triyas stratigrafisi (Palaeozoic and Triassic stratigraphy of Gondwanan Platform). Proceedings A. Suat Erk Symposium on Geology, Ankara University Publication, Ankara, 3–9.Google Scholar
d'Orbigny, A. 1826. Tableau méthodique de la clssa de Céphalopodes. Annales des Sciences Naturelles 7, 245–314.Google Scholar
Dunbar, C. O. & Henbest, L. G. 1930. The fusulinid genera Fusulina, Fusulinella and Wedekindella. American Journal of Science, Series 5, 20, 357–64.CrossRefGoogle Scholar
Dunbar, C. O. & Skinner, J. W. 1931. New fusulinid genera from the Permian of West Texas. American Journal of Science, Series 5 22, 252–68.CrossRefGoogle Scholar
Dunbar, C. O. & Skinner, J. W. 1936. Schwagerina versus Pseudoschwagerina and Paraschwagerina. Journal of Paleontology 10, 83–91.Google Scholar
Epshteyn, O. G., Terekhova, T. P. & Solovieva, M. N. 1985. Paleozoic of Koryak Range (Foraminiferal fauna, Biostratigraphy). Voprosy Mikropaleontologii 27, 47–77.Google Scholar
Forke, H. C. 1995. Biostratigraphie (Fusuliniden, Conodonten) und Mikrofazies im Unterperm (Sakmar) der Karnischen Alpen (Nassfeldgebiet, Österreich). Jahrbuch der Geologischen Bundesanstalt 138, 207–97.Google Scholar
Forke, H. C. 2002. Biostratigraphic subdivison and correlation of Uppermost Carboniferous/Lower Permian sediments in the Southern Alps: Fusulinoidean and conodont faunas from the Carnic Alps (Austria/Italy), Karavanke Mountains (Slovenia) and Southern Urals (Russia). Facies 47, 201–76.CrossRefGoogle Scholar
Fursenko, A. V. 1958. Fundamental state of development of foraminiferal faunas in the geological past. Trudy Instituta Geologicheskikh Nauk, Akademiia Nauk Belorusskoi SSR, Minsk 1, 10–29.Google Scholar
Groves, J. R. 2000. Suborder Lagenina and other smaller foraminifers from uppermost Pennsylvanian–lower Permian rocks of Kansas and Oklahoma. Micropaleontology 46, 285–326.Google Scholar
Groves, J. R. & Boardman, D. R. 1999. Calcareous smaller foraminifers from the Lower Permian Council Grove Group near Hooser, Kansas. Journal of Foraminiferal Research 29, 243–62.Google Scholar
Gürçay, B. 2000. Aydıncık (İçel) batisinin jeolojisi (Geology of western Aydıncık (İçel). MTA Cumhuriyetin 75. Yıldonumu Yerbilimleri ve Madencilik Kongresi Bildiriler Kitabi I. MTA Publication, Ankara, 93–105.Google Scholar
Gutnic, M., Monod, O., Poisson, A. & Dumont, J. F. 1979. Géologie des Taurides Occidentales (Turquie). Mémoires de la Société Géologique de France 137, 1–109.Google Scholar
Güvenç, T. 1977 a. Stratigraphie du Carbonifère et du Permien de la Nappe de Hadim. In 6th Colloquium on Geology of the Aegean Regions (eds Izdar, E. & Nakoman, E.), pp. 251–61. Aegean University, Izmir.Google Scholar
Güvenç, T. 1977 b. Permian of Turkey. In 6th Colloquium on Geology of the Aegean Regions (eds Izdar, E. & Nakoman, E.), pp. 263–82. Aegean University, Izmir.Google Scholar
Güvenç, T. 1980. Alanya-Gazipaşa bolgesinin jeolojisi ve kiyiseridi deniz tabaniyla yapisal iliskileri (Geology of Alanya-Gazipaşa area and the structural relations with sea-bed). Aegean University Institute of Marine Sciences and Technology, Izmir, 139 pp.Google Scholar
Güvenç, T. 1991. Stratigraphy and paleontology of Permian in Anatolia. International Congress on Permian System of the World, 1991, Part 1, Perm, Russia. Occasional Publications, Earth Science and Research Institute, New Series 8B, 29–39.Google Scholar
Güvenç, T., Demirel, I. H., Meshur, M., Gül, M. A. & Tekin, U. K. 1991. The paleogeography of Anatolia during the Permian and Triassic. Occasional Publications, Earth Science and Research Institute, New Series 11B, Part III, 11–40.Google Scholar
Hanzawa, S. 1939. Stratigraphical distribution of the genus Pseudoschwagerina and Paraschwagerina in Japan with description of two new species of Pseudoschwagerina from Kitami Mountainland, Northeastern Japan. Japan Journal of Geology and Geography 16 (1–2), 65–73.Google Scholar
Hanzawa, S. 1961. Facies and Micro-Organism of the Paleozoic, Mesozoic and Cenozoic sediments of Japan and its adjacent islands. International Sedimentology and Petrography, Series 5, 117 pp.Google Scholar
Kahler, F. 1976. Die Fusuliniden der Dorud Formation im Djadjerud-Tal nordlich von Tehran (Iran). Rivista Italiana di Paleontologia e Stratigrafia 82, 439–66.Google Scholar
Kahler, F. & Kahler, G. 1937. Beiträge zur Kenntnis der Fusuliniden der Ostalpen: Die Pseudoschwagerinen der Grenzlandbänke und des Oberen Schwagerinenkalkes. Palaeontographica, Abteilung A 87, 1–43.Google Scholar
Kahler, F. & Kahler, G. 1966. Fossilium Catalogus I; Animalia Fusulinida (Foraminiferida) parts 1–4, 934 pp. s’-Gravenhage (Junk).Google Scholar
Kahler, F. & Kahler, G. 1980. Fusuliniden aus den Kalken des Trogkofel-Schichten der Karnischen Alpen. Die Trogkofel-Stufe im Unterperm der Karnischen Alpen. Carinthia II 36, 183–254.Google Scholar
Kalmykova, M. A. 1967. Permian fusulinids of Darvaz. Transactions, VSEGEI 116 (2), 116–285.Google Scholar
Kanmera, K., Ishii, K. & Toriyama, R. 1976. The evolution and extinction patterns of Permian fusulinaceans. Geology and Paleontology of Southeast Asia 17, 129–54.Google Scholar
Kanmera, K. & Mikami, T. 1965. Fusuline zonation of the Lower Permian Sakamotozawa Series. Memoirs of the Faculty of Science, Kyushu University, Serie D, Geology 16, 275–320.CrossRefGoogle Scholar
Krainer, K. & Davydov, V. I. 1998. Facies and biostratigraphy of the late Carboniferous/Lower Permian sedimentary sequence in the Carnic Alps (Austria/Italy). Geodiversitas 20, 643–62.Google Scholar
Lee, J. S. 1927. Fusulinidae of North China. Palaeontologica Sinica, Series B 4, 1–172.Google Scholar
Lefevre, R. 1967. Un nouvel élément de la géologie du Taurus Lycien: Les nappes d'Antalya (Turquie). Comptes Rendus de l'Academie des Sciences, 7, série D 265, 1365–8.Google Scholar
Leven, E. Ja. 1971. Les gisements permiens et les Fusulinidés de l'Afghanistan du Nord. Notes et Mémoires Moyen-Orient 12, 1–46.Google Scholar
Leven, E. Ja. 1975. Stage-scale of the Permian deposits of Tethys. Moscow Society of Naturalists, Geological Series, Bulletin 50, 5–21.Google Scholar
Leven, E. Ja. 1993. Early Permian fusulinids from the Central Pamir. Rivista Italiana di Paleontologia e Stratigrafia 99, 151–98.Google Scholar
Leven, E. Ja. 1995. Lower Permian fusulinids from the vicinity of Ankara. Rivista Italiana di Paleontologia e Stratigrafia 101, 235–48.Google Scholar
Leven, E. Ja. 1997. Permian stratigraphy and fusulinida of Afghanistan with their paleogeographic and paleotectonic implications. Geological Society of America Special Paper 316, 1–134.Google Scholar
Leven, E. Ja. & Davydov, V. I. 2001. Stratigraphy and fusulinids of the Kasimovian and Lower Gzhelian (Upper Carboniferous) in the southwestern Darvaz (Pamir). Rivista Italiana di Paleontologia e Stratigrafia 107, 3–46.Google Scholar
Leven, E. Ja. & Okay, A. 1996. Foraminifera from the Exotic Permo-Carboniferous Limestone Blocks in the Karakaya Complex, Northwestern Turkey. Rivista Italiana di Paleontologia e Stratigrafia 102, 139–74.Google Scholar
Leven, E. Ja. & Shcherbovich, S. F. 1978. Fusulinids and stratigraphy of the Asselian stage of Darvaz. Akademiya Nauk SSSR, Nauka Publishing House, 1–162.Google Scholar
Leven, E. Ja. & Shcherbovich, S. F. 1980. Sakmarian fusulinid assemblage of Darvaz. Voprosy Mikropaleontologii 23, 71–85.Google Scholar
Lin, J. X., Li, J. X., Chen, G. X., Zhou, Z. R. & Zhang, B. F. 1977. Fusulinida. In Paleontological Atlas of Central South China 2, (Late Paleozoic) (ed. Hubei Institute of Geological Sciences), pp. 4–96.Google Scholar
Marcoux, J. 1976. Le Trias des nappes à radiolarites et ophiolites d'Antalya (Turquie): Homologies et signification probable. Bulletin de la Societe Géologique de France 18, 511–12.CrossRefGoogle Scholar
Miklukho-Maklay, A. D. 1949. Verkhnepaleozoyskie fuzulinidy Sredney Azii, Fergana, Darvaz i Pamir. Leningradskiy Gosudarstvennyy Universitet, 111 pp.Google Scholar
Miklukho-Maklay, A. D. 1959. Znachenie gomeomorfii dlya sistematiki fuzulinid. Uchenye Zapiski Leningradskogo Gosudarstvennogo Universiteta 268, Seriya Geologicheskogo Nauk 10, 155–72.Google Scholar
Moeller, V. von. 1878. Die spiral-gewundenen Foraminiferen des russischen Kohlenkalks. Mémoires de l'Académie Impériale de Sciences de St. Petersbourg, Sér. 7 25, 1–147.Google Scholar
Morikawa, R. & Isomi, H. 1960. A new genus Biwaella, Schwagerina-like Schubertella. Science Reports of the Saitama University, Urawa, Series B 3 (3), 301–5.Google Scholar
Morikawa, R. & Isomi, H. 1961. Studies of Permian Fusulinids in the East of Lake Biwa, Central Japan. Geological Survey of Japan, Report 191, 1–29.Google Scholar
Ota, Y. 1997. Middle Carboniferous and Lower Permian fusulinaceans biostratigraphy of the Akiyoshi Limestone Group, Southwest Japan. Part I. Bulletin of the Kitakyushu Museum of Natural History 16, 1–97.Google Scholar
Ota, Y. 1998. Middle Carboniferous to Early Permian fusulinaceans from the Akiyoshi Limestone Group, Southwest Japan. Bulletin of the Kitakyushu Museum of Natural History 17, 1–105.Google Scholar
Ozawa, T., Kobayashi, F., Ishii, Ken-Ichi, & Okimura, Y. 1990. Carboniferous to Permian Akiyoshi Limestone Group. Fourth International Symposium on Benthic Foraminifera, Guidebook for Field Trip 4, E1–E31.Google Scholar
Ozawa, T., Kobayashi, F. & Watanabe, K. 1991. Biostratigraphic zonation of Late Carboniferous to Early Permian sequence of the Akiyoshi Limestone Group, Japan and its correlation with reference sections in the Tethyan region. Proceedings of the Shallow Tethys 3, Saito Ho-on Special Publication, Sendai, Japan, 327–41.Google Scholar
Özgül, N. 1976. Toroslarin bazi temel jeolojik ozellikleri (Some geological aspects of the Taurus orogenic belt (Turkey)). Bulletin of the Geological Society of Turkey 19, 65–78.Google Scholar
Özgül, N. 1984. Stratigraphy and tectonic evolution of the Central Taurides. In Geology of the Taurus Belt, International Symposium (eds Göncüoğlu, M. C. & Tekeli, O.), pp. 77–90. Ankara: General Directorate of Mineral Research and Exploration (MTA).Google Scholar
Putrya, F. S. 1956. Stratigraphy and foraminifera of Middle Carboniferous strata of the eastern Donbass. Trudy Vsesoyuznogo Neftyanogo Nauchno-issledovatel'skogo Geologo-razvedochnogo Instituta (VNIGRI), Leningrad, n. s. 98, 33–485.Google Scholar
Rauser-Chernousova, D. M. 1937. Rugosofusulina, a new fusulinid genus. Etyudy po Mikropaleontologii, Paleontologicheskaya Laboratoriya Moskovskogo Gosudarstvennogo Universiteta, Moscva 1, 9–26.Google Scholar
Rauser-Chernousova, D. M., Bensh, F. R., Vdovenko, M. V., Gibshman, N. B., Leven, E. Ya., Lipina, O. A., Reitlinger, E. A., Solovieva, M. N. & Chediya, I. O. 1996. Spravochnik po sistematike foraminifer paleozoya (endothyroidy, fusulinoidy). Rossiiskaya Akademiya Nauk, Geologicheskii Institut, Nauka, 207 pp.Google Scholar
Rozovskaya, S. E. 1975. Composition, phylogeny and system of the order Fusulinida. Trudy Paleontologicheskogo Instituta, Akademiya Nauk SSSR 149, 267 pp.Google Scholar
Sanders, D. & Krainer, K. 2005. Taphonomy of Early Permian benthic assemblages (Carnic Alps, Austria): carbonate dissolution versus biogenic carbonate precipitation. Facies 51, 539–57.CrossRefGoogle Scholar
Schellwien, E. 1898. Die Fauna des Karnischen Fusulinidenkalks. Teil II. Palaeontographica (1897) 44, 237–82.Google Scholar
Schellwien, E. 1908. Monographie der Fusulinen. Teil I, Die Fusulinen des Russisch-Arktischen Meeresgebietes. Palaeontographica 55, 145–94.Google Scholar
Shcherbovich, S. F. 1969. Fusulinids of the late Gzhelian and Asselian time of the Precaspian Syneclise. Trudy Geologicheskogo Instituta, Akademiya Nauk SSSR 176, 1–82.Google Scholar
Shlykova, T. I. 1948. Fusulinids from the Upper Carboniferous of the Samara Bend. In Microfauna of the oil fields of the USSR, pp. 1–131. VNIGRI, Sbornik 1, Trudy, n. s., vypusk 31.Google Scholar
Skinner, J. W. 1931. Primitive fusulinids of the Mid-Continent region. Journal of Paleontology 5, 253–9.Google Scholar
Skinner, J. W. & Wilde, G. L. 1954. Fusulinid wall structure. Journal of Paleontology 28, 445–51.Google Scholar
Staff, H. Von & Wedekind, R. 1910. Der oberkarbonische Foraminiferen-Sapropelit Spitzbergens. Bulletin of the Geological Institution of the University of Uppsala 10, 81–123.Google Scholar
Suleimanov, I. S. 1949. New species of Fusulinidae of the Subfamily Schubertellinae Skinner from the Carboniferous and Lower Permian of the Bashkirian Cis-Urals. Trudy Geologicheskogo Instituta Akademiya Nauk 105, Geologicheskaya Seriya 35, 22–43.Google Scholar
Tekeli, O. 1980. Toroslarda Aladağlarin yapisal evrimi (Structural evolution of Aladağ Mountains in Taurus Belt). Bulletin of the Geological Society of Turkey 23, 11–14.Google Scholar
Tekeli, O., Aksay, A., Urgun, B. M. & Işik, A. 1984. Geology of the Aladağ mountains. In Geology of the Taurus Belt, International Symposium (eds Göncüoğlu, M. C. & Tekeli, O.), pp. 143–58. Ankara: General Directorate of Mineral Research and Exploration (MTA).Google Scholar
Toriyama, R. 1958. Geology of Akiyoshi. Part 3. Fusulinids of Akiyoshi. Memoirs of the Faculty of Science, Kyushu University, ser. D, Geology 7, 1–264.CrossRefGoogle Scholar
Ulakoğlu, S. 1983/1984. Aladağlarda Yahyalı (Kayseri) bolgesinin jeolojisi (The geology of Yahyalı regions (Kayseri) in the Aladağ Mountains. Istanbul University Engineering Faculty's Earth Sciences Review 4, 1–44.Google Scholar
Vachard, D. 1990. A new biozonation of the limestone from Terbat area (Sarawak, Malaysia). In Ten years of CCOP research on the Pre-Tertiary of East Asia (ed. Fontaine, H.), pp. 193–208. Economic and Social Commission for Asia and the Pacific, Committee for Co-ordination of Joint Prospecting for Mineral Resources in Asian Offshore Areas (CCOP), Technical Publication no. 20.Google Scholar
Vachard, D. & Krainer, K. 2001. Smaller foraminifers characteristic algae and pseudo-algae of the Latest Carboniferous–Early Permian Rattendorf Group, Carnic Alps (Austria/Italy). Rivista Italiana di Paleontologia e Stratigrafia 107, 169–95.Google Scholar
Wang, L. W. 1974. A handbook of the stratigraphy and paleontology in southwest China. Nanjing Institute of Geology and Paleontology, Academia Sinica, Science Press, 454 pp.Google Scholar
Watanabe, K. 1991. Fusuline biostratigraphy of the Upper Carboniferous and Lower Permian of Japan, with special reference to the Carboniferous–Permian boundary. Palaeontological Society of Japan Special Papers 32, 1–150.Google Scholar
White, M. P. 1932. Some Texas Fusulinidae. Texas University Bulletin 3211, 1–104.Google Scholar
Wu, O., Lin, C. & Wu, Y. 1986. Late Carboniferous and Latest Permian fusulinid faunas from Jingshe, Longyan District of Fujian and their stratigraphic significance. Acta Micropalaeontologica Sinica 3 (1), 13–32.Google Scholar
Yabe, H. 1938. Carboniferous–Permian deposits of the Japanese Islands, Tyosen (Korea) and Manchuria. Compte Rendu, 2nd International Carboniferous Stratigraphy Congress (Heerlen, 1935), 1617–49.Google Scholar
Zhao, Z., Han, J. X. & Wang, Z. 1984. The Carboniferous strata and its fauna from southwestern margin of Tarim Basin in Xijiang. Geological Publishing House, pp. 1–154.Google Scholar
Zhou, J. P. 1991. Fusulinid zones from Maping Formation of Changmo, Longlin, Guangxi; on Carboniferous–Permian boundary. Acta Palaeontologica Sinica 30, 396–405.Google Scholar
Zhou, T. M., Sheng, J. H. & Wang, Y. J. 1987. Carboniferous–Permian boundary beds and fusulinid zones at Xiaodushan Guangnan; Eastern Yunnan. Acta Micropalaeontologica Sinica 4, 123–60.Google Scholar
Zhou, Z. 1982. Earliest Permian Schwagerina cushmani fusulinids from southeastern Hunan. Acta Palaeontologica Sinica 21, 225–48.Google Scholar