High-precision U-Pb zircon ages on SE Newfoundland tuffs now bracket the Avalonian Lower–Middle Cambrian boundary. Upper Lower Cambrian Brigus Formation tuffs yield depositional ages of 507.91 ± 0.07 Ma (Callavia broeggeri Zone) and 507.67 ± 0.08 Ma and 507.21 ± 0.13 Ma (Morocconus-Condylopyge eli Assemblage interval). Lower Middle Cambrian Chamberlain’s Brook Formation tuffs have depositional ages of 506.34 ± 0.21 Ma (Kiskinella cristata Zone) and 506.25 ± 0.07 Ma (Eccaparadoxides bennetti Zone). The composite unconformity separating the Brigus and Chamberlain’s Brook formations is constrained between these ages. An Avalonian Lower–Middle Cambrian boundary between 507.2 ± 0.1 and 506.3 ± 0.2 Ma is consistent with maximum depositional age constraints from southwest Laurentia, which indicate an age for the base of the Miaolingian Series, as locally interpreted, of ≤ 506.6 ± 0.3 Ma. The Miaolingian Series’ base is interpreted as correlative within ≤ 0.3 ± 0.3 Ma between Cambrian palaeocontinents, although its exact synchrony is questionable due to taxonomic problems with a possible Oryctocephalus indicus-plexus, invariable dysoxic lithofacies control of O. indicus and diachronous occurrence of O. indicus in temporally distinct δ 13C chemozones in South China and SW Laurentia. The lowest occurrence of O. indicus assemblages is linked to onlap (epeirogenic or eustatic) of dysoxic facies. A united Avalonia is shown by late Early Cambrian volcanics in SW New Brunswick; Cape Breton Island; SE Newfoundland; and the Wrekin area, England. The new U-Pb ages revise Avalonian geological evolution as they show rapid epeirogenic changes through depositional sequences 4a–6.

Postel Nunatak in the Patuxent Range has been previously mapped as Nelson Limestone but there was no biostratigraphic support for that interpretation until now. We confirm that limestone exposures at Postel Nunatak are at least partly correlated with the Nelson Limestone of the Neptune Range, 160 km north-east, and are not correlative with the lower Cambrian Schneider Hills Limestone of the Argentina Range. Upper beds have yielded the trilobites Suludella? davnii Palmer & Gatehouse, 1972 and Solenopleura pruina Palmer & Gatehouse, 1972, which provide a basis for assignment to Cambrian Series 3 (late middle Cambrian), within the Drumian or lower Guzhangian stages. Limestone beds were deposited in a shallow marine setting, ranging from supratidal to lagoonal facies with rare subtidal intervals. These settings contrast with deeper water facies of the Neptune Range. Despite limitations in sampling density, isotopic analysis indicates that a greater than +2.5‰ shift in δ13C is consistent with δ13C trends documented for the Drumian Stage. Because the upper and lower contacts at Postel Nunatak are covered by snow and ice, the relationship with rocks mapped as the Patuxent Formation in the Patuxent Range remains uncertain, but part of it may belong to the Precambrian Hannah Ridge Formation.

Diverse and well-preserved acritarchs are reported from the type section of the Cambrian Hanford Brook Formation at Hanford Brook, southern New Brunswick. This section fills an important gap in acritarch studies by providing the first detailed picture of changing acritarch associations close to the traditional lower–middle Cambrian boundary in Avalonia. Acritarchs from the St Martins Member, at the base of the succession, include Skiagia ciliosa, Heliosphaeridium notatum, H. longum and Liepaina plana and suggest attribution to Cambrian Stage 4. Acritarchs from the Somerset Street Member, in the middle of the formation, include Eliasum llaniscum and Comasphaeridium silesiense. This information adds new biochronological context to an ash bed in the Somerset Street Member previously dated as c. 510 Ma or 508 Ma, and to the endemic trilobites from the same member, including Protolenus elegans. It also places absolute ages on the basal range of stratigraphically important acritarchs. Both the acritarch assemblage and the radiometric age are consistent with a position very close to the traditional lower–middle Cambrian transition and likely within Cambrian Stage 5. Acritarchs from the Long Island Member, at the top of the succession, include additional taxa demonstrating assignment to Cambrian Stage 5. Both the Somerset Street and Long Island members probably correlate with the Morocconus notabilis Zone. The new acritarch species Retisphaeridium striatum Palacios is described. New data are presented on acritarchs from the upper part of the Hell's Mouth Formation, Wales, and correlation proposed with the Long Island Member.

A volcanic tuff 1.0 m above the base of the Triebenreuth Formation in the Franconian Forest provides the first precise and biostratigraphically bracketed date within the traditional Middle Cambrian. The first illustration of fossils from the Triebenreuth Formation in this report and their discussion allow a more highly refined correlation within the Middle Cambrian. A weighted mean 206Pb–238U date of 503.14±0.13/0.25/0.59 Ma on zircons from this subaerial pyroclastic tuff was determined by U–Pb chemical abrasion isotope dilution mass spectrometry (CA-TIMS) techniques. At c. 6.0–7.0 Ma younger than the base of the traditional Middle Cambrian in Avalonia, the new West Gondwanan date from east-central Germany suggests that estimates of 500 Ma for the base of the traditional Upper Cambrian and 497 Ma on the base of the Furongian Series may prove to be too ‘old’. Biostratigraphically well-bracketed dates through most of the Middle Cambrian/Series 3 and below the upper Upper Cambrian/upper Furongian Series do not exist. An earlier determined 494.4±3.8 Ma date from the Southwell Group of Tasmania may actually prove to be a reasonable estimate for the age of the base of the traditional Upper Cambrian. Until high precision dates are determined on the base of the traditional Upper Cambrian and base of the Furongian Series, the rates of biotic replacements and geological developments and the durations of biotic zones in the Middle/Series 3 and Upper Cambrian/Furongian Series remain as ‘best guesses’.

A revision of the trilobite fauna from Brzechów in the Holy Cross Mountains (Poland) shows that the assemblage consists of Palaeolenus medius (Bednarczyk), Issafeniella orlowinensis (Samsonowicz), Kingaspidoides sanctacrucensis (Czarnocki), Paradoxides (Acadoparadoxides) oelandicus Sjögren, P. (A.) cf. mureroensis Sdzuy and P. (A.) sp. It is earliest Middle Cambrian in age according to the West Gondwana standard of Geyer & Landing (2004) and can be correlated with the Cephalopyge notabilis Zone of Morocco. Moreover, the co-occurrence of the ranges of P. (A.) oelandicus with P. (A.) cf. mureroensis might suggest that P. (A.) oelandicus appears earlier in the Brzechów sandstones than in Scandinavia, thus the strata exposed here do not find their correlative in the Scandinavian succession. As a result, the Banian/Agdzian boundary (Lower–Middle Cambrian boundary according to the West Gondwana standard) probably lies within the Kamieniec Formation, and also below the Brzechów sandstones, in the upper part of the Ociesęki Formation. Close similarity of the discussed assemblage with contemporaneous assemblages from Morocco, Jordan or New Brunswick indicates that the fauna is generally of peri-Gondwanan and Gondwanan aspect.

Paradoxides jemtlandicus Wiman, 1903, from strata near the boundary between the oelandicus Stage and the paradoxissimus Stage of the middle Cambrian in central Sweden, is redescribed using more complete specimens than were known hitherto. It is one of the earliest species of the genus Paradoxides sensu stricto, and though the form of the dorsal exoskeleton and the transverse shape of the hypostome are normal for the genus, compared with typical Paradoxides such as P. paradoxissimus it shows features that can be regarded as primitive. These features include the shape of the glabella, the lack of strong fusion of the rostrum with the hypostome, the acute inner spine angle of the librigena and the short pygidium; they are reminiscent of some species of Hydrocephalus.

The middle Cambrian Lejopyge laevigata Zone is poorly exposed in Scandinavia. Both this zone, however, and the succeeding Agnostus pisiformis Zone are well exposed at a classic locality at Gudhem, Västergötland, south-central Sweden. The sequences consist of finely laminated alum shale with scattered stinkstone (orsten) lenses. Three measured and sampled sections yielded a diverse fossil fauna, dominated by trilobites, in particular agnostoids, and the bradoriid Anabarochilina primordialis. Fossils are excellently preserved but restricted to the stinkstones. The L. laevigata Zone at Gudhem includes several geographically widespread key agnostoid species, notably Tomagnostella sulcifera, Clavagnostus spinosus, Glaberagnostus altaicus, Lejopyge laevigata and L. armata. The L. laevigata Zone in Scandinavia is here extended to include the traditional Solenopleura? brachymetopa Zone, and its lower boundary is defined by the FAD of L. laevigata. Trilobite evidence shows that the upper part of the Scandinavian L. laevigata Zone approximately correlates with the Proagnostus bulbus Zone of China and elsewhere.

The modes of life of twenty three species of arthropods from the Shale are reviewed, with special reference to locomotion and feeding. Six groups are recognised, predatory and scavenging benthos, deposit-feeding benthos, scavenging and possibly predatory nektobenthos, deposit-feeding and scavenging nektobenthos, nektonic filter-feeders, and a miscellaneous sixth group that includes a questionable example of parasitism and a species that doubtfully grazed on algae. These animals had but limited powers of walking, digging, raking or swimming. Within these limitations a range of morphological adaptations and modes of feeding had been evolved, by Middle Cambrian time, a range comparable to that found in Recent marine forms. Arthropods in the Shale dominated, in numbers of individuals and possibly in biomass, a fauna dwelling above, on and in a muddy substrate at a depth of about 100 m.