The Punta Peligro fauna includes some of the oldest Cenozoic South American mammals, and the oldest ones for Patagonia. In addition to frogs, turtles, and crocodiles, an unusual mammalian assemblage is formed by a mixture of Mesozoic lineages of Gondwanan origin and therians (eutherians and metatherians) derived from Laurasian immigrants. This paper describes new remains of the Didolodontidae ‘condylarth’ genus Escribania Bonaparte, Van Valen & Kramarz, 1993. The new material includes an isolated right lower third molar with talonid morphology different from the homologous structure of the type species Escribania chubutensis Bonaparte, Van Valen & Kramarz, 1993, justifying the recognition of a new species. The derived nature of the didolodontid ‘condylarths’ from the Banco Negro Inferior, their differences with the Mioclaenidae Kollpaniinae from the early Paleocene of Tiupampa, and the record of bona fide litoptherns in the Banco Negro Inferior (BNI) suggest an early radiation of the panameriungulates.

A total of 47 genera, in 80 species, of Mississippian, or Lower Carboniferous, crinoids are evaluated from 61 localities in Scotland based on a modern update of the literature, study of museum collections, and new field work. Among the 80 species, 76 are considered valid, with eight requiring new combinations of genus and species names. In addition, one species is considered a nomen dubium, and three can only be assigned at the generic level. Asbian faunas have a moderate generic richness of 13. Brigantian faunas have the highest generic richness at 40. Arnsbergian faunas have the lowest generic richness at three. There are no Mississippian crinoid faunas in Scotland older than Asbian. The dominant Brigantian crinoid faunas occur in limestones and shales in the shallow shelf marine intervals of Yoredale-style cyclothems. The Asbian faunas occur in shallower, nearshore mudstones and limestones of Yoredale-style cyclothems. The late Viséan (Brigantian) fauna is more similar in overall taxonomic composition to the global record for the Serpukhovian rather than the Viséan. This reflects the establishment of the Late Palaeozoic Crinoid Macroevolutionary Fauna, dominated by advanced cladid crinoids, by the end of the Viséan in Scotland.

A study of type and newly collected material of the Silurian ostracod Richteria migrans (Barrande, 1872) demonstrates that it had wide distribution, occurring in at least the Czech Republic, France, Sardinia, Wales, central Asia and probably Poland. R. migrans has biostratigraphic value, as an indicator for the mid to late part of the Ludlow Series. It was almost certainly pelagic, living predominantly in probable shelf topographic lows to marginal/off-shelf environments, characteristically with cephalopod-graptolite-bivalve-dominant associates. It had at least five to six growth stages, exhibits polymorphic variation, and its morphology provides evidence to endorse the notion that ‘entomozoaceans’ are myodocopes.

The Late Carboniferous whip spider Graeophonus anglicus Pocock, 1911 (Arachnida: Amblypygi), is redescribed on the basis of the holotype and nine other specimens all preserved in sideritic nodules from the British Middle Coal Measures of Coseley, Staffordshire, UK. This species is clearly basal with respect to most living whip spiders, expressing numerous plesiomorphic character states and can be referred to both the suborder Paleoamblypygi and the ‘living fossil’ family Paracharontidae (with one Recent species), the latter based on an explict character of dorsal spination on the pedipalp femur. This suggests that crown-group Amblypygi originated by at least the mid-Palaeozoic.

New cranial material of the ‘rhynchodipterid’ lungfish Soederberghia groenlandica from Upper Devonian (Famennian) strata in East Greenland is described. Previously unknown structures identified here include components of the hyoid arch (ceratohyal, hypohyal) and the lower jaw (prearticular, dentary). Earlier interpretations of the cheek and mandible of Soederberghia are reconsidered in the light of new fossil specimens. Some of the difficulties in assessing the homologies between cheek bones in Soederberghia and those of other lungfishes stem from confusion over the arrangement in Rhynchodipterus, and a revised interpretation of this genus is proposed. The single infraorbital bar found in Soederberghia probably originates, in part, from an expanded bone 10 (quadratojugal) of the kind found in Griphognathus. Hypotheses that posit ‘rhynchodipterid’ polyphyly seem unlikely in light of a set of derived cranial characters that define a coherent radiation of long-snouted, denticle-bearing lungfishes known from the Late Devonian. The hypothesis presented here places ‘rhynchodipterids’ as a paraphyletic grade with respect to fleurantiids. Rhynchodipterus, Soederberghia, and fleurantiids form a clade to the exclusion of the species of Griphognathus. G. minutidens is the sister taxon to this apical group, while G. sculpta and G. whitei are more remote from it.

Dark limestones in the old quarries at Leny, Perthshire contain sparse beds with tiny fossils. They are poorly preserved and, though barely affected by the Ordovician Grampian Event tectonism, there is some taphonomic distortion and many are corroded along stylolitised horizons. The fauna mainly comprises trilobites of two types, open-ocean miomerids and polymerid shelf dwellers. Miomerids Condylopyge cf. eli and Kiskinella cristata indicate a stratigraphical position equivalent to the base of the paradoxidid Amgan Stage of Siberia; traditionally regarded as ‘Middle Cambrian’. However, the bulk of the Leny miomerids, notably species of Pagetides, are forms described from the outer edge of Laurentia, within the Bonnia–Olenellus Zone, where it is considered to be ‘Lower Cambrian’. The Leny polymerids were likely transported off-shelf and some are conspecific with taxa in the Laurentian allochthonous Quebec and New York successions of the Early–Middle Ordovician (Taconic) Appalachian Orogen. The Leny Limestone and Shale Member of the Keltie Water Grit Formation is part of the Dalradian Supergroup deposited in an off-shelf Caledonide Grampian Terrane of the Humber Tectonostratigraphical Zone, midway between the North American successions and the Greenland Caledonides.

Additional to the trilobites, brachiopods, sponges, hyoliths, bradoriids and a selection of indeterminable organic fragments occur; none of which has any particular age significance.