The Permian–Triassic climate crisis can provide key insights into the potential impact of horizon threats to modern-day biodiversity. This crisis coincides with the same extensive environmental changes that threaten modern marine ecosystems (i.e., thermal stress, deoxygenation and ocean acidification), but the primary drivers of extinction are currently unknown. To understand which factors caused extinctions, we conducted a data analysis to quantify the relationship (anomalies, state-shifts and trends) between geochemical proxies and the fossil record at the most intensively studied locality for this event, the Meishan section, China. We found that δ18Oapatite (paleotemperature proxy) and δ114/110Cd (primary productivity proxy) best explain changes in species diversity and species composition in Meishan’s paleoequatorial setting. These findings suggest that the physiological stresses induced by ocean warming and nutrient availability played a predominant role in driving equatorial marine extinctions during the Permian–Triassic event. This research enhances our understanding of the interplay between environmental changes and extinction dynamics during a past climate crisis, presenting an outlook for extinction threats in the worst-case “Shared Socioeconomic Pathways (SSP5–8.5)” scenario.

The common observation that smaller particle-size fractions of sedimentary rocks yield younger K-Ar apparent ages than the larger particle-size fractions of the same stratigraphic age was analyzed with the aid of the 40Ar/40K ratio from 14 stratigraphically and regionally different sections. Estimation of the loss of radiogenic 40Ar from varied clay-rich size fractions was based on two models: a relationship between particle size and the 40Ar/40K ratio, and a theoretical diffusional loss from spherical particles. The differences between the two models and reconciliation of their results are discussed. For the smallest fractions (up to <0.5 μm), percent-wise losses of 40Ar from the spherical particles model increase from Upper Carboniferous and Permian (38±10%), to Late Triassic (47±10%), and to Miocene and Late Neogene (65±8%). This trend suggests that escape of 40Ar from the smaller particles in older sediments decreased or even stopped after deposition of the sedimentary sections.

The large 40Ar losses derived from small 40Ar/40K ratios in the younger Tertiary sediments, indicate that addition of K to the small fractions is, at least in part, responsible for the young K-Ar apparent ages in geologically different settings. In several 102–103 m thick sections, authigenic illite in the <0.1 to <2 μm fractions yields young K-Ar apparent ages resulting from simultaneous 40Ar production and release during clay authigenesis. In a production and loss model, a first-order escape-rate parameter (e) was estimated at 0.2 × 10−8 to 4 × 10−8 y−1, depending on the K-Ar apparent age of the size fractions and the stratigraphic age of the section. The limitations and uncertainties of the methods of evaluating diagenetic 40Ar losses from fine clay particles are discussed.

Diagenetic kaolin minerals are very common in the Permo-Triassic succession from the SE Iberian Range, Spain. The morphology and crystal structure of kaolin minerals has been examined in four size fractions (<1 µm, <2 µm, <6.3 µm and <20 µm) of sandstone samples by means of scanning electron microscopy, X-ray diffraction, infrared spectroscopy, differential thermal analysis and thermogravimetry. Experimental data reveal that dickite is the dominant kaolin-type mineral in the entire range of size fractions, whereas small amounts of kaolinite coexists with dickite in all size fractions. Dickite appears typically as booklets of pseudo-hexagonal plates with blocky habit. The increase in size fraction is concomitant with the increase in the amount of dickite and the progressive improvement of its structural order. The extensive dickitization is attributed to the high paleogeothermal gradient recorded in the studied area and the increase in H+, presumably resulting from the flux of organic acids derived from the underlying Carboniferous rocks and/or the late Permian succession. These conditions are more likely to be associated with the late Cretaceous post-rift thermal stage of the eastern Iberian Basin. Lately, during the maximum burial depth, the fine crystalline kaolin minerals were slightly illitized. Given the very small feldspar content in the studied sequence, the results reflect the important contribution of mica alteration to the early diagenetic formation of kaolinite as well as the late conversion to dickite.

The Upper Permian sedimentary successions in the northern Sydney Basin have been the subject of several stratigraphic, sedimentological and coal petrographic studies, and recently, extensive U-Pb zircon dating has been carried out on tuffs in the Newcastle Coal Measures. However, detailed petrographic and geochemical studies of these successions are lacking. These are important because a major change in tectonic setting occurred prior to the Late Permian because of the Hunter-Bowen Orogeny that caused the uplift of the Carboniferous and Devonian successions in the Tamworth Group and Tablelands Complex adjacent to the Sydney Basin. This should be reflected in the detrital makeup of the Upper Permian rocks. This study provides data that confirms major changes did take place at this time. Petrographic analysis indicates that the source area is composed of sedimentary, felsic volcanic and plutonic and low-grade metamorphic rocks. Conglomerate clast composition analysis confirms these results, revealing a source region that is composed of felsic volcanics, cherts, mudstones and sandstones. Geochemical analysis suggests that the sediments are geochemically mature and have undergone a moderate degree of weathering. The provenance data presented in this paper indicate that the southern New England Orogen is the principal source of detritus in the basin. Discrimination diagrams confirm that the source rocks derive from an arc-related, contractional setting and agree with the provenance analyses that indicate sediment deposition in a retroarc foreland basin. This study offers new insights on the provenance and tectonic setting of the Northern Sydney Basin, eastern Australia.

Pareiasaurs were globally distributed, abundant, herbivorous parareptiles with the basal-most members found only in the mid-Permian of South Africa. These basal forms form a monophyletic group and were locally abundant and became extinct at the top of the Tapinocephalus Assemblage Zone at the end of the Guadalupian. Four species of basal pareiasaurs are currently recognised: Bradysaurus baini, B. seeleyi, Embrithosaurus schwarzi and Nochelesaurus alexanderi, but they are all poorly understood and there remains historic uncertainty as to their validity. In this paper, our second contribution designed to improve understanding of the basal group, we present the first detailed cranial description and updated diagnosis for Nochelesaurus alexanderi and demonstrate that it is a distinct taxon based on one cranial autapomorphy, a large transversely wide postparietal, and a combination of cranial characters. Within the local group of mid-Permian pareiasaurs, we recognise new dental features of Nochelesaurus alexanderi: non-symmetrical marginal cusp arrangements on upper and lower teeth resulting from an extra basal mesial cusp; an incipient horizontal cingulum on lower jaw teeth, sometimes with one or two tiny medial cingular cusps; and up to ten marginal cusps. Our study demonstrates that tooth morphology and orientation, cranial ornamentation, morphology of the cheek bosses, shape of the postfrontal and postparietal, and morphology of the distal paroccipital process of the opisthotic are the most useful to identify South African mid-Permian pareiasaurs.

This paper is devoted to the X-ray diffraction study of bone fragments of Permian parareptile Deltavjatia vjatkensis obtained from the Kotelnich vertebrate fossil site, one of the richest of the Permian period, which is characterized by the excellent preservation of fossil remains because of their burial in a silty anaerobic environment similar to modern bogs. The bone is well-preserved and consists of fluorapatite, calcite, quartz, and dolomite. The refined apatite unit-cell parameters of a and b-axis (9.3526 ± 0.0001 and 9.3587 ± 0.0001) Å and c-axis (6.8930 ± 0.0001 and 6.8968 ± 0.0001) Å correspond to F-apatite. Crystallinity index determined as the full width at half maximum of the 002 reflection in degrees 2θ is 0.266–0.250, which is typical for Mesozoic vertebrate bones. Apatite crystallite size (length 70.3–74.9 nm, width 30.7–30.3 nm) in fossil pareiasaur bone is larger than in subfossil and recent mammal bone and is in a good agreement with the values for seismosaurus bone. Both crystallite size and aspect ratio (2.3–2.5) are independent of the fossil pareiasaur bone length.

Several types of authigenic illite differing in their distribution and morphology are described in Rotliegend (Permian) gas-bearing sandstones of northern Germany. The microtextural environment seems to control the K-Ar age variations of illite-fractions of different sizes. Thin, uncompacted, and radial coatings are 200 Ma old, whereas uncompacted fibrous laths filling enlarged secondary porosity are 180 Ma old, and compacted or recrystallized tangential particles yield intermediate or even younger ages. Such Liassic ages relate to a thermal event, the effects of which on the clay material have not been erased subsequently, despite present-day burial depth of 4500 m and temperatures of 150°C.

The nature, distribution and origin of clay minerals in the hydrocarbon-bearing Permian Rotliegend sandstones of the North Sea and the adjacent areas of the Netherlands and Germany are reviewed. The clay minerals occur as detrital coatings of smectite and smectite-illite on the surfaces of sandgrains, and as later diagenetic cements of kaolinite, chlorite (two varieties), and illite in the pore spaces of those sandstones. Two diagenetic clay mineral assemblages are predominant in the Rotliegend of the North Sea. The kaolinite-illite assemblage is restricted to the Rotliegend of shelf areas which underwent shallow burial followed by strong Jurassic/Cretaceous (Late Cimmerian) structural inversions, whereas the illite-chlorite assemblage is associated with basinal areas that underwent deep and rapid burial throughout the Mesozoic.

The factors controlling mineralogy, crystal chemistry and morphology of those diagenetic clay minerals, as well as their regional distribution and origin, are numerous, complicated, and inter- related. Evidence suggests that the following aspects were important parameters: (1) variations in the original depositional arid desert environment; (2) the chemistry and flow patterns of the porewaters; (3) temperature and timing of clay mineral formation; (4) local burial history; (5) the presence or absence of meteoric water; and (6) the structural setting of the Rotliegend sandstones.

Oxygen isotope data indicate that the illite cements formed over a wide range of temperatures (24–140°C) that is consistent with the deep burial conditions prevailing in the palaeo-basins. In contrast, oxygen isotopes indicate that kaolinite cements formed over a more restricted temperature range (40–80°C) and under the influence of meteoric water penetrating the sandstones of the shelf areas as a result of their Late Cimmerian uplift and associated erosion. Hypotheses suggesting that the absence of kaolinite cement from the deeply buried Rotliegend sandstones is caused by its illitization during burial, and that the chlorite cements have formed by the alteration of earlier smectite, smectite-chlorite and corrensite cements, are not supported by evidence.

Detailed structural, volcanic, and sedimentary investigations of the crustal response to the emplacement of the Middle–Late Permian Emeishan large igneous province show that a rifting system developed prior to the main stage of flood basalt eruptions, in the form of sedimentary breccias, clastic sedimentary deposits and mafic hydromagmatic units. Detrital zircon grains from sandstones yield ~750–800 Ma LA-ICP-MS 206Pb/238U age clusters, showing that material was sourced from the Yangtze crystalline basement. Gabbros and pegmatites intruded along the normal faults of the rift system yield zircon ages of 264–260 Ma, and thus constrain the timing of rifting. N–S-trending rift zones developed along the western flank of the Pan-Xi palaeo-uplift, with NE–SW- and NNE–SSW-trending rifts on the eastern side and along the western and eastern margins of the Yangtze Block. The rifting progressed in pulses, with an initial phase of normal faulting followed by rapid deposition of breccias. Later there was lower-energy deposition of sandstone, with accompanying rhyolitic eruptions. This was followed by low-energy sedimentation of mudstones and dolomites, with accompanying hydromagmatic deposits. Rift system formation was constrained by a combination of far- and near-field tectonic stresses due to plate motions and lithospheric interaction with initial Emeishan volcanism.

Sudoite, an aluminium-rich di-trioctahedral chlorite, is known primarily from highgrade diagenetic/ low-temperature metamorphic Al-rich rocks and from hydrothermal deposits in Alrich terrains. This contribution reports a rare occurrence of pore-lining sudoite in Permian red beds from the eastern part of the Southern Permian Basin. Sudoite is a ubiquitous mineral in aeolian sandstones of the Eastern Erg, a large dune field buried in the subsurface of the Fore-Sudetic Monocline, SW Poland. Sudoite crystals are arranged in a honeycomb texture and often associated with high-porosity intervals. It is concluded that sudoite crystallized at the expense of early diagenetic dioctahedral smectite at a temperature lower than 180°C under the influence of hot, Mgrich and K-poor fluids, which migrated from a basement and/or deep part of the basin upward through permeable aeolian sandstones and mixed with colder pore solutions. Tosudite was a likely intermediate phase in the smectite to sudoite conversion.

The granitic rocks of the Tarim large igneous province (LIP) are temporally and spatially related to mafic intrusions. However, their tectonic setting and genetic relationship are debated. Here, we report geochemical, and zircon U–Pb–Hf isotopic results for three alkali feldspar granitic plutons in the Halajun area, western margin of the Tarim Block. Zircon U–Pb ages suggest these plutons were emplaced at 268–275 Ma, coeval with the neighbouring mafic–ultramafic complexes and syenitic rocks. These granitic rocks have high contents of SiO2, alkalis, Rb, Th, Zr and REE (except Eu), and high ratios of FeO*/MgO and Ga/Al, and show strong depletions in Ba, Sr, Eu, which are commonly observed in the A1-type granites. Zircon Hf isotopes reveal a limited range of εHf(t) values from −1.0 to +3.5 for different samples from three granitic plutons, obviously higher than those (mostly <0) of the mafic rocks. This distinct difference, along with a Daly gap and small volume of mafic rocks, argues against extreme fractionation of mafic magma as the main origin of the A1-type granites. Instead the A1-type granites were most likely derived from partial melting of newly underplated rocks triggered by the upwelling asthenosphere, followed by extensive fractionation. These A1-type granites were emplaced within an anorogenic setting during the late stage of the Tarim LIP, which possibly lasts for more than 30 Ma. The Piqiang mafic–ultramafic complex directly stemmed from asthenospheric mantle and Halajun A1-type granites represent two manners of vertical crustal growth.

A reconnaissance survey of part of western Cambodia has found quarries in and near the town of Pailin containing chert, basalt, volcaniclastic rocks, gabbro and serpentinite. Abandoned quarries contain chert and basalt clasts in a volcaniclastic matrix and may constitute a mélange. A new, recently active, quarry contains these rock units in close and coherent contact but they lack the serpentinitic matrix of the Thung Kabin Mélange of eastern Thailand. The rock units are in close contact and may best be described collectively as a dismembered ophiolite. The mélange–ophiolite association constitutes a 3 km wide, 20 km long E–W belt separating a northern 200 km2 block of the mainly amphibolitic Pailin Crystalline Complex from a southern area of Triassic submarine fan siliciclastic rocks. The cherts yield the first documented radiolarian fauna from Cambodia and include a moderately well-preserved Asselian–Sakmarian age fauna consisting of Pseudoalbaillella sakmarensis, Pseudoalbaillella scalprata morphotype scalprata, Pseudoalbaillella sp. cf. P. simplex, Pseudoalbaillella u-forma morphotype II, Pseudoalbaillella sp. cf. P. elegans, Pseudoalbaillella sp. cf. P. lomentaria, Albaillella sp., Pseudoalbaillella spp., Trilonche? sp., Latentifistularia gen. et sp. indet. and Entactinaria gen. et sp. indet. The Pailin ophiolitic rocks, mélange and volcanic rocks occur within a generally E–W-trending belt, which suggests that the Sa Kaeo Suture does not extend southeastwards paralleling the Thai–Cambodian border, nor extend under the Cardamom Mountains but, rather, extends eastwards into Cambodia and possibly then turns southwards along the strike of the Pursat–Kampot Fold Belt.

Late Permian fluvio-lacustrine successions of the Çakraz Formation in the Zonguldak Terrane between the regions of Akçakoca and Ereğli were investigated in order to describe the litho- and biostratigraphic properties and explain the depositional environment. The studied succession with black, dark-grey to greenish-grey shales, siltstones and limestones is named the Alaplı Member to distinguish it from the classical red clastic successions, which are tentatively named the Ereğli Member of the Çakraz Formation. The organic-rich black shales, mudstones and limestones of the Alaplı Member yielded palynological assemblages suggesting a Lopingian (Tatarian) age. The lack of any marine macro- or microfossils, the fine-grained character of the lithofacies with abundant plant material and the association of poorly sorted conglomerates in the middle part of the succession indicate possible deposition in a broad range of fluvial and lacustrine environments. Successions of similar age and depositional environment are known from the East European Variscan Belt in Bulgaria and Romania. Common successions were also developed in actively extending shallow-marine platforms on the NW Palaeotethyan margin at the end of the Permian Period.

Since the beginning of the development of the North German Basin in Stephanien to Early Rotliegend times, rifting played a major role. Nearly all structures in NW-Germany and the German North Sea - (more than 800) - salt diapirs, grabens, inverted grabens and inversion structures - are genetically related to rifting. Today, the rifting periods are well dated. We find signs of dilatation at all times except from the Late Aptian to the end of the Turonian. To the contrary, the period of the Coniacian and Santonian, lasting only five million years was a time of compression, transpression, crustal shortening and inversion. Rifting activities decreased notably after inversion in Late Cretaceous times. Tertiary movements concentrated on a limited number of major, long existing lineaments. Seismically today NW-Germany and the German North Sea sector is one of the quietest regions in Central Europe.

This paper presents depth maps for eight key horizons and seven thickness maps covering the onshore and offshore areas for the Late Permian to recent sedimentary section of the Netherlands. These maps, prepared in the context of a TNO regional mapping project, are supported by nine regional structural cross sections and a table summarizing the timing of tectonic activity from Carboniferous to recent. These new regional maps enable the delineation of various structural elements but also reveal the development of these elements through time with improved detail. Since the latest Carboniferous the tectonic setting of the Netherlands changed repeatedly. During successive tectonic phases several pre-existing structural elements were reactivated and new elements appeared. The various identified regional structural elements are grouped into six tectonically active periods: Late Carboniferous, Permian, Triassic, Late Jurassic, Late Cretaceous and Cenozoic. This study demonstrates that many structural elements and fault systems were repeatedly reactivated and that a clear distinction exists between long-lived elements, such as the Roer Valley Graben, and short-lived structural elements, such as the Terschelling Basin.

Chert and silicified wood from the Permian through Cretaceous of Antarctica contain abundant information on fungal diversity and plant–fungal interactions. The chert deposits represent a particularly interesting setting for the study of plant–fungal interactions because they preserve remains of distinctive high latitude forest ecosystems with polar light regimes that underwent a profound climate change from icehouse to greenhouse conditions. Moreover, some of the cherts and wood show the predominance of extinct groups of seed plants (e.g. Glossopteridales, Corystospermales). Over the past 30 years, documentation of fossil fungi from Antarctica has shifted from a by-product of plant descriptive studies to a focused research effort. This paper critically reviews the published record of fungi and fungal associations and interactions in the late Palaeozoic and Mesozoic cherts and silicified wood from Antarctica; certain fungal palynomorphs and fungal remains associated with adpression fossils and cuticles are also considered. Evidence of mutualistic (mycorrhizal), parasitic and saprotrophic fungi associated with plant roots, stems, leaves and reproductive organs is presented, together with fungi occurring within the peat matrix and animal–fungus interactions. Special attention is paid to the morphology of the fungi, their systematic position and features that can be used to infer fungal nutritional modes.

Two crinoid pluricolumnals from Permian rocks of Timor show similar patterns of external pitting. Platycrinitid sp. preserves circular, parabolic pits that do not cross-cut between columnals, some have raised rims and at least one columnal shows a growth deformity. These pits are interpreted as the trace fossil Oichnus paraboloides Bromley. Crinoid sp. indet. has particularly dense pits cross-cutting columnals on one side of the pluricolumnal only and extending onto the contiguous limestone; it is a Holocene microkarstic solution feature. Care must be taken to separate true bioerosive trace fossils from modern microkarstic features in limestones in the tropics.

The end-Permian (c. 252 Ma) and end-Triassic (c. 201 Ma) mass-extinction events are commonly linked to the emplacement of the large igneous provinces of the Siberia Traps and Central Atlantic Magmatic Province, respectively. Accordingly, scenarios for both extinctions are increasingly convergent and cross-fertilization of ideas has become important. Here, we present a synthesis of extinction scenarios based on a critical assessment of the available palaeontological, sedimentological, geochemical and geophysical evidence. How similar were the extinction events, what gaps exist in our understanding and how can a comparison of the events enhance our understanding of each event individually? Our focus is on the most important proximate kill mechanisms including: climate change and atmospheric pollution; increased soil erosion, weathering and runoff; forest dieback and the spread of pathogens; and ocean temperature changes, anoxia and acidification. There is substantial evidence to suggest that very similar kill mechanisms acted upon late Permian as well as Late Triassic ecosystems, strengthening the hypothesis that the ultimate causes of the mass-extinction events were similar.

New material of the pareiasaur Deltavjatia rossicus from the Kotel'nich locality, Kirov Province, Russia, is described in detail. The taxon is characterised by a distinctive pattern of dermal sculpture and the exaggerated embayment of the posterior skull roof, resulting in the dorsal exposure of the braincase. Postcranially, Deltavjatia shares some aspects of its morphology with basal pareiasaurs, including the osteoderm pattern. Features such as the forward-slanting and pointed iliac blade are shared with stratigraphically younger, more derived forms. Well-preserved material of the taxon spans a wide size-range, allowing an assessment of ontogenetic trends. A geometric morphometric analysis of the skull roof of Deltavjatia reveals an allometric increase in snout length and postorbital area, a result that can serve as a basis for examining morphological trends within pareiasaurs. A reassessment of pareiasauromorph relationships, using both parsimony and Bayesian methods of phylogenetic inference, recovers similar topologies in both cases. Four Bayesian analyses were completed, with and without a gamma-shaped parameter and with and without the inclusion of autapomorphies. Despite differing taxon and outgroup selection, the recovered topologies are similar to previous phylogenies of pareiasaurian relationships, with Deltavjatia appearing as a relatively basal taxon.

New geochronological data for the Haolibao porphyry Mo–Cu deposit, NE China, yield Permian crystallization zircon U–Pb ages of 278 ± 5 Ma for granite and 267 ± 10 Ma for the granite porphyry that hosts the Mo–Cu mineralization, and four Re–Os molybdenite ages yield an isochron age of 265 ± 3 Ma. These ages disagree with the previous K–Ar age determinations that suggest a correlation of intrusive rocks of the Haolibao area with the Yanshanian intrusive rocks of Cretaceous age. The mineralizations at the Haolibao area may be related to the tectonic–magmatic activity caused by collisional events between the North China Plate and Mongolian terranes during the Permian. The occurrence of the Haolibao plutonic rocks indicates that the Palaeo-Asian-Mongolian Ocean closed during the Permian along the Xilamulun River suture.