Freeliving corals capable of automobility (e.g., lateral migration) were rare during Paleozoic time, but some species within the tabulate genera Procterodictyum, Procteria (Granulidictyum), P. (Pachyprocteria), Palaeacis and Smythina, and the rugose genera Combophyllum, and Baryphyllum, have morphologic characters that suggest they were capable of such self-directed movement. The rugose corals Gymnophyllum and Hadrophyllum, sensu stricto may have exhumed and righted themselves. No single morphological character is diagnostic for an automobile habit, but the following characters appear to be important indicators: 1) lack of an external attachment surface; 2) concentric skeletal accretion; 3) discoid corallum shape; 4) concavo-convex, plano-convex, and, less commonly, biconvex corallum profile; and 5) small, lightweight corallum. Additionally, the occurrence of corallites on the base of the corallum (hypocorallites) is a good indicator of automobility in freeliving corals, but the character is so far known only in Procterodictyum. All known fossil automobile taxa appear to have inhabited relatively quiet environments on muddy or silty, soft substrates.

The earliest known automobile corals were early Emsian (Devonian) Procterodictyum. Paleozoic automobile corals were most abundant during Devonian time, with Procterodictyum, Procteria (Granulidictyum), and Combophyllum distributed in a narrow longitudinal band in the southern hemisphere on both sides of the Rheic Ocean. Carboniferous and Permian automobile taxa (Palaeacis partim, Smythina and Baryphyllum) were less diverse, but more cosmopolitan. Throughout Paleozoic time, the vast majority of automobile corals was confined to within 40 degrees of the paleoequator. However, additional research will be required before coral automobility can be used to constrain paleolatitude independently.

Taxonomy of heliolitine corals, and tabulate corals in general, benefits from analysis of larger assemblages of coralla that approximate populations. Quantitative data derived from such assemblages allow intraspecific variation to be tested rigorously and illustrated, and similar, co-occurring species to be discriminated using normal patterns of continuous population variation. Field collection and analysis necessarily involve recording and comparing assemblages from the full spectrum of sedimentary facies available at any one stratigraphic level and from the maximum available stratigraphic range. The large published taxonomic literature on heliolitine corals is a daunting obstacle to identification at the species level because a substantial part appears to involve taxonomic splitting to a degree inconsistent with variation known in modern coral species. The heliolitine corals studied occur in Ludlow-age ramp and shelf carbonate facies in the Boothia Uplift region in the southern Arctic archipelago of Canada. The “population” approach resulted in discrimination of 18 morphotaxonomic species, belonging to five genera and four families. A few select morphologic characters, incorporated in simple identification keys, are particularly helpful in distinguishing the 11 heliolitid and four stelliporellid species present, and are useful in screening the voluminous literature on species of Heliolites. Six of the species compare closely to Silurian species reported from central Kazakhstan, West Siberia, and North China; few are shared with other regions. The principal affinities are consistent with occurrence in the Uralian-Cordilleran paleogeographic region in the Silurian northern hemisphere. Eight species do not appear to be recorded in published literature elsewhere and are apparently endemic.

The current classification of scleractinian corals based upon gross morphological features has been found unsatisfactory due to additional information from skeletal microarchitecture and microstructure. It is necessary to investigate microstructural details and limits in morphologic variation within and between different coral clades before a revised classification is constructed. Variations in morphologic characters and microstructural details from a population of Dimorpharaea de Fromentel, 1861 (Family Microsolenidae) from Upper Bathonian (Jumara Dome) strata in Kachchh are described. The data used include the diameter (D) and height (H) of the corallum, number of corallites in the colony (NC), number of septa in the mother corallite at the center of the colony (NS), minimum distance between centers of central corallite and corallite of the inner ring (C1), minimum distance between corallite centers of the outer ring (C2), septal density (DS) and trabecular density (DT). The principal components analysis reveals that most of the variation is explained by “size” related characters (D and H) while corallite density (NC and C1) and septal structures (DS and DT) contribute to the second and third principal component axes, respectively. The microarchitecture and distribution of characters observed in the Kachchh Dimorpharaea require a re-evaluation of familial-specific concepts and suggest that the population belongs to a single species, Dimorpharaea stellans Gregory, 1900, rather than four nominal species (D. stellans, D. distincta, D. continua and D. orbica) as has been assumed.

Extremely well-preserved specimens of the species Rennensismilia complanata and Aulosmilia cuneiformis occur in Santonian (Upper Cretaceous) strata of the Lower Gosau beds, near Gosau, Austria. Two of these, here reported, have aragonitic skeletal mineralogy and skeletal structures that are typical for their families, and, in addition, show distribution of trace elements (Sr and Mg above all) that confirm the biogenic origin of structures observed. R. complanata also has proteinaceous matrix preserved within its skeleton, which, seen here for the first time in electron micrographs of living or fossil corals, forms sheaths of organic matrix surrounding bundles of skeletal crystallites. Matrix is most abundant along the axial plane of septa, which also is the first-formed part of each septum. Although A. cuneiformis lacks observable organic matrix materials, its skeletal structure and its distribution and amount of trace elements are analogous to that seen in R. complanata and also in modern corals.

Dicoelosia occurs in two deep water benthic shelly assemblages on an Early Silurian (uppermost Aeronian, Stimulograptus sedgwickii Zone) carbonate ramp to shelf, within the 25 m thick bluish-grey mudstone of the Richardson Member in the middle Jupiter Formation, Anticosti Island, Québec. Dicoelosia dauphinensis new species is erected on the basis of its relatively large, elongate shell, with a concavo-convex lateral profile, moderately wide, planoconvex lobes and subparallel lateral margins. Dicoelosia dauphinensis first occurs in a Gotatrypa-Dicoelosia Community, in which it constitutes about 7 percent of the individuals within sampled populations. This is replaced about 2 m up section by a Resserella-Dicoelosia Community, where D. dauphinensis makes up 22 percent of the individuals preserved on single bedding planes. Subsequently, Dicoelosia becomes a rare component of the Stegerhynchus-Triplesia Community in a shallowing-upward succession of the upper Richardson Member. It is in this upper Richardson shelly community that Stimulograptus sedgwickii is locally common, together with in situ brachiopod nests of Eocoelia, Triplesia, and Lissatrypa at the upper Jupiter Cliff section. The water depth estimated for the Resserella-Dicoelosia Community is between 100-120 m, below the normal depth range of the Clorinda Community (BA5) on Anticosti, in a distal shelf setting about 80-100 km offshore from the Laurentia paleocontinent, on the west side of the Iapetus Ocean. The interpretation of water depth is based on the facts that, 1) Dicoelosia-rich communities are confined to strata, which lack shallow water sedimentological evidence such as thick calcarenites and hummocky cross stratification, 2) Dicoelosia does not occur with shallower water taxa seen in the overlying Clorinda and Stricklandia communities associated with cyclocrinitid algae, corals and stromatoporoids in the Cybèle Member, and 3) Dicoelosia occurs only in the deeper water sections of the Richardson Member, some 10-15 km basinwards from mid-shelf shallower Richardson facies of the Anticosti Basin. Thus, the Dicoelosia-rich communities are interpreted to mark a maximum flooding surface within the distal shelf to ramp Llandovery succession of Anticosti Island.

The component subfamilies of the Delthyridoidea are critically reviewed and subjected to phylogenetic analysis. This shows the presence of two clades, assigned to the Delthyrididae and Acrospiriferidae, within the superfamily. The subfamilial categories are redefined mainly on the basis of the characters used in the phylogenetic analysis. The spiriferid, mainly delthyridide, Gaspé fauna is formally revised and redescribed. This new taxonomic treatment leads to more precise biostratigraphy and to the recognition of a new subfamily, the Gaspespiriferinae, based on the new genus Gaspespirifer. Five new species are described: Howellella (Howellella) forillonensis, Brachyspirifer (Brachyspirifer) briseboisi, Paraspirifer desbiensi, Brevispirifer florentinus, and B. quebecensis, The occurrence of Brevispirifer species with Middle Devonian chonetaceans confirms the presence of marine Eifelian strata in the Matapédia Valley. Paraspirifer desbiensi n. sp. and two species left in open nomenclature, Vandercammenina sp. and Mucrospirifer sp., have considerable biostratigraphic and biogeographic significance in the Lower Devonian. The first occurrence in the Eastern Americas Realm of the typically Rhenish genera Brachyspirifer, Paraspirifer, and Vandercammenina are in Gaspé. This reinforces the hypothesis that Gaspé served as a stepping stone for Rhenish species invading North America in Pragian and Emsian times, as previously suggested by bivalve biogeography.

The only two known life associations of the shell and operculum of Maclurites Le Sueur, 1818, are described for the first time. Maclurites cf. magnus Le Sueur, 1818, from the Middle Ordovician of Alaska demonstrates that the operculum fits slightly inside the aperture, and when closed, the operculum extends down below the base of the shell and may have acted as an anchor. The operculum of Maclurites bigsbyi (Hall, 1861), from the Middle Ordovician of Wisconsin, is illustrated for the first time and is confirmed to be distinct from Maclurina logani (Salter, 1859).

Gastropods are described for the first time from the Kersadiou Formation (middle Givetian) in the vicinity of Brest, Brittany, northwestern France. Twenty-three species of gastropods are recognized. New taxa include the new genera, Calvibembexia, Kersadiella, Breizospira, Finisterella, Parahormotomina, and Nodoloxonema, represented by the new species Calvibembexia lethiersi, Kersadiella babini, Breizospira crozonensis, Finisterella tibidiensis, Parahormotomina sibertae, and Nodoloxonema plusquelleci. Further new species include: Sinuitina (Sinuitina) morzadeci, Crenistriella armoricana, Patellilabia (Phragmosphaera) ponceti, Angyomphalus (Eoangyomphalus) weyanti, Eoplatyzona ongaroae, Murchisonia oehlerti, and Hormotomina fiacrensis. The biogeographic affinities of the fauna show a strikingly unusual admixture of Eastern Americas Realm taxa such as Hormotomina and Crenistriella. This suggests that during the middle Givetian, the Massif Armoricain (northwestern France) represented an area of Realm-boundary mixing between the Old World and Eastern Americas Realms, while older Devonian gastropod faunas of the Massif Armoricain are wholly of Old World Realm type.

The Goodland/Comanche Peak Limestone, Kiamichi Formation and basal Duck Creek Limestones around Fort Worth Texas yield a limited number of cosmopolitan ammonite and inoceramid bivalve taxa that allow precise correlation with the sequence that has been used as a standard in northwest Europe. The upper part of the Goodland/Comanche Peak Limestones yields species of Dipoloceras that show the base of the Upper Albian substage, provisionally defined as the first appearance of D. cristatum (Brongniart, 1822), to lie within this unit. Brancoceras aff. cricki Spath, 1934, Mortoniceras (Deiradoceras) beloventer new species, and Actinoceramus cf. concentricus (Parkinson, 1819) parabolicus Crampton, 1996a, co-occurs with D. cristatum in the Comanche Peak Limestone. The Kiamichi Formation yields rare Mortoniceras (Mortoniceras) pricei (Spath, 1922), M. (Deiradoceras) prerostratum Spath, 1921, M. (D.) bipunctatum Spath, 1933, and Actinoceramus sulcatus (Parkinson, 1819) morphotypes that allow correlation with the European Hysteroceras orbignyi and Hysteroceras varicosum subzones of the Mortoniceras inflatum zone. The basal Duck Creek Limestone yields Mortoniceras (Deiradoceras) sp. and Hysteroceras cf. varicosum (J. de C. Sowerby, 1824), and can also be correlated with the varicosum subzone.

Alokistocare subcoronatum (Hall and Whitfield, 1877) is the type species of Alokistocare Lorenz, 1906, the nominal genus for the diverse Middle Cambrian Alokistocaridae Resser, 1939b. New collections from the Ehmaniella Biozone contain cranidia, librigenae, hypostomes, rostral plates, thoracic segments, and pygidia, which allow a more complete understanding of the morphological range and character of this important species.

Cladistic analyses using this new information about A. subcoronatum suggest that: 1) Ehmaniellidae Sundberg, 1994, is synonymous with Alokistocaridae; 2) Alokistocaridae is a monophylectic clade and not synonymous with Papyriaspididae Whitehouse, 1939; 3) Pseudoalokistocare Sundberg, 1994, is synonymous with Alokistocare; and 4) Taxa previously assigned to the older “Alokistocare”/Amecephalus group (Plagiura to Glossopleura biochrons) form a separate clade and should not be included in Alokistocare or the Alokistocaridae. These older species are assigned to Amecephalus Walcott, 1924, which is restricted to the Delamaran Stage. Alokistocaridae as recognized here is restricted to the Marjuman Stage.

A morphometric analysis of Alokistocare subcoronatum shows minor but significant differences among collections, indicating population differences. In addition, the cranidium displays both positive and negative allometric growth as well as isometric growth. Allometric growth is present in both meraspid and holaspid stages. The pygidium shows only isometric growth.

Pliomerina Chugaeva, 1958, a characteristic element of the peri-Gondwanan Eokosovopeltis-Pliomerina Province, occurs in Llanvirn and Caradoc strata of the Argentine Precordillera. Three protaspid instars are present in the ontogeny of the Early Caradoc Pliomerina peripata new species. A species possibly representing Pliomerella is a rare element in Early Caradoc strata of the Las Aguaditas Formation. Prosopiscus is represented by a Llanvirn species with a first record of protaspid larvae for the enigmatic Family Prosopiscidae. Early growth stages support closest relations of prosopiscids to Phacopina. The presence of Pliomerina and Prosopiscus, along with the cheirurid Pateraspis, demonstrates a peri-Gondwanan influence on the Precordilleran fauna by the Llanvirn, and supports the proximity of the Precordillera terrane to the South American margin by that time.

New cheirurid trilobite species from the Las Aguaditas Formation in San Juan Province include the Llanvirn Macrogrammus rafi, and the Early Caradoc Ceraurinella zhoui, Nieszkowskia jakei, and N. yongyii. Precordilleran species, such as the Llanvirn Macrogrammus pengi new species from the Gualcamayo Formation, reinforce sinoparapilekiinid affinities for the poorly known Macrogrammus Whittard, 1966, although the monophyly of Sinoparapilekiini requires corroboration. The ontogeny of Ceraurinella zhoui confirms the presence of early protaspid stages for this genus (and Cheirurinae in general). Rare cheirurids include species of Heliomeroides, Pateraspis, and an additional acanthoparyphinid.

An isolated premaxillary tooth of a tyrannosaurid from the Lower Cretaceous section of the Tetori Group, Central Honshu, Japan, complements Siamotyrannus, which is based upon an incomplete postcranium for existence of tyrannosaurids in the Early Cretaceous of Asia. The occurrence of a tyrannosaurid tooth in the Japanese early Early Cretaceous further supports the possibility that tyrannosaurids originated during the Early Cretaceous in Asia and migrated to North America when the two continents were connected via a land bridge during the early Late Cretaceous. Thickening of the premaxillary teeth might have predated the increase in body size in tyrannosaurid evolution.

A new agriochoerid, Protoreodon walshi new species, from the middle Eocene Santiago Formation, San Diego County, California, is described from the Jeff's Discovery and Rancho del Oro local faunas, considered to be late Uintan in age. It differs from most other species of Protoreodon in having an undivided P4 para-metacone and a P3 protocone connected to the paracone by a buccolingual crest. Associated postcranial material is abundant and well preserved. The postcranial skeleton shows several features which probably represent the primitive oreodont condition, including closely appressed metapodials, hoofed ungual phalanges, the lunar in broad contact with both the magnum and the unciform in anterior view, and having the magnum larger than the trapezoid.

Two specimens of a larger morph, Protoreodon cf. walshi, show a different P3 morphology and might represent another new species. Diversity of agriochoerid oreodonts in California is at least as high as has been reported for the Vieja Group of Texas, although not as high as in the Uinta Basin of Utah, where the genus Diplobunops is also present.

Two new species, Mojavemys galushai and Phelosaccomys neomexicanus, are described from the Barstovian of New Mexico. Both of these species are assigned to a new subfamily, the Mojavemyinae. It is uncertain as to whether this new subfamily should be referred to the Heteromyidae or Geomyidae.

Both Mojavemys an Phelosaccomys retain some heteromyid characteristics while developing advanced traits of geomyids. It is possible that these genera represent transitional forms between a heteromyid ancestor and advanced geomyines, thus deriving the Geomyidae directly from the Heteromyidae sometime in the middle Tertiary. It is also possible that the Mojavemyinae are a subfamily of heteromyids only convergent with geomyids. A third possiblity is that the mojavemyines are among a group of middle Tertiary geomyoids from which both Recent heteromyids and geomyids were derived.

Phosphatic microfossils including probable microvertebrates occur in the Early and Middle Ordovician rocks of Wisconsin. The eight types described here include the probable ostracoderm Anatolepis, and six unidentified but possible vertebrate fragments as well as the palaeoscolecid worm Milaculum. These fossils are rare, but at two localities several specimens per kg were recovered over significant stratigraphic intervals. The high concentration of certain fossils in the lower part of the Galena Group suggests that there is stratigraphic value in their description. Identification of similar material or of articulated assemblages of these fossils may eventually lead to identification of biologic affinities.