The chapter explores the background to Churchill’s upbringing and its bearing on his future career. It introduces and contrasts his American Jerome lineage with his aristocratic British Marlborough ancestry before describing the courtship and early married life of Jennie Jerome and Lord Randolph Churchill. Winston’s education is placed in the context of the meteoric rise and sudden collapse of his father’s political career, the family’s financial difficulties and Lord Randolph’s failing health. The chapter finishes with an analysis of Churchill’s family inheritance and concludes that its importance was more social and political than financial.

This chapter traces the history of the Tolstoy family, dating Lev Nikolayevich Tolstoy’s ancestry back to the fourteenth century. It begins with a brief explanation of the Tolstoy family’s complicated genealogy, distinguishing between the titled branch and the untitled Tver and Kutuzov branches. Next there is a brief overview of the Tolstoy family’s most famous scions, ranging from prominent statesmen to flamboyant individuals whose colorful lives consolidated the family reputation for eccentricity. There is also a brief discussion of the distinguished and troubled history of the Volkonsky family, Tolstoy’s maternal ancestors. The chapter then focuses on Tolstoy’s immediate family – his parents, brothers, sister, aunts and, of course, his wife Sofia Andreevna and their children. There is commentary on what they meant to him, both personally and creatively, and on how he drew on them, as well as his more distant forebears, to breathe flesh and blood into the iconic characters of his two great novels.

The chapter opens with a concise history of the recognition of blood types that, as well as informing the human immune system, incidentally provide an ideal genetic data set for phylogenetic enquiry. The chapter goes on to relate that enquiry to the author’s People of the British Isles project. This project draws upon genetic data from just over 2,000 volunteers to construct a genetic map of the human population of the British Isles. From that genetic map, the peopling of the British Isles is inferred and reconstructed.

This chapter breaks with the typical detachment of chroniclers of Russian politics from the social wavelengths that in so many ways defy conventional periodization. Stalin may have proclaimed a classless society, and inconvenient memories were “frozen” out of discourse, yet social distinctions continued to be cognitively programmed among future generations in families and in the realm of the professional, educational, and social institutions to which they were exposed. First, I discuss theorizing on historical memory. This helps us distinguish between articulated injunctions and situational staging on the one hand and unarticulated silences on the other, as expressed in choices, symbols, or artifacts. Next, I present survey results, which corroborate not only hypothesized covariance between educated estate ancestry and white-collar occupations but, intriguingly, awareness of the placement of respondents’ forbears in the imperial estate structure. In the remaining sections, I make sense of these patterns by discussing our protagonists’ journey of ancestral reconstruction; tapping into reminiscences of private spaces and public-oriented practices within them; exploring familial injunctions and artifacts that create mental cartographies and ties structuring exclusion and belonging; and, finally, discerning the descendants’ instrumentalizations of ancestry to actively delineate their place in post-communist Russian society.

When individuals sign on to a DNA ancestry test, they understand that the company will undertake an analysis of certain segments of their genome, called ancestry information markers (AIMs). These segments can, under proper analysis, reveal their genetic descent from certain regions of the world.

Over a period of 20 years, family genetic genealogy, through the purchase of consumer ancestry testing kits, has been one of the fastest growing family activities of this generation. Citing data from the International Society of Genetic Genealogy, the Washington Post reported in 2017 that eight million people worldwide were involved with recreational genomics. It is estimated that by 2019 about 25 million people had signed up for a DNA ancestry test offered by one of the dozens of companies that have entered this marketplace. The kits are sent to a person’s home with return packaging that includes a reservoir for depositing saliva or swabs for sampling cheek cells. The MIT Technology Review predicted that by 2021 there would be 100 million consumers of ancestry DNA services.

Most human genetic diversity is found within populations rather than between populations. Scientists have reported that any two individuals within a particular population are as different genetically as any two people selected from any two populations in the world. Given this finding, how can science use a small percentage of genetic diversity between populations as markers of ancestral origins?

Much of recreational DNA ancestry offers consumers a long reach into the history of their descent by discovering which biogeographical population most closely matches their DNA profiles. The science and DNA analytics provide probability estimates that their DNA markers (ancestry informative markers, or AIMs) are most likely from a particular continent or even a specific country. But DNA ancestry tests have applications that go well beyond recreational genealogy. Even prior to the growth of this sector of direct-to-consumer testing, DNA was used to determine paternity and to establish identity in criminal investigations. An important and largely unintended application of ancestry DNA testing has been the uncovering of family secrets: “Why does my father look so different from his parents?” or “Why are my mother’s skin tones so much darker than those of her parents?”

As we noted previously, the science behind DNA ancestry requires that one compares the unique genetic markers on the consumer’s DNA sample with the frequency of those markers in reference panels representing different regions of the world. When the field of DNA ancestry began, it was a scientific project that involved the search for biogeographical DNA. Scientists could use changes in the human genome to determine how ancient populations moved around the globe. The further populations moved across the globe and the more time elapsed (many thousands of years), the greater the number of mutations or genetic variants. Genetic ancestry began with a half-dozen distinct continental regions and with markers called hypervariable microsatellites, or short tandem repeats (STRs) of DNA, 2–6 base pairs in length. These microsatellites were considered ideal at the time because they had a high heterozygosity, which means two different alleles at a site. A site that has an AA is homozygous, whereas one that has AG is heterozygous. The more diverse the alleles, the greater the chance of distinguishing allele frequencies among populations. Initially, scientists used changes in the maternally inherited mitochondrial DNA (mtDNA) and the paternally inherited Y chromosome. That changed when autosomal markers were chosen for ancestry analysis.

In order to locate people’s ancestry to a region of the world through their DNA, the markers on their DNA sample have to be compared to population reference panels for the regions that form part of the comparison group. These ancestry inference methods have served medical research, forensic science, and commercial popular genealogical interests. According to Santo et al., the reliability of any ancestry inference depends on the existence of reliable population reference databases. Many researchers and ancestry DNA companies utilize different sources for population data on different countries. For example, ALFRED is an allele frequency database supported by the Yale Center for Medical Informatics, which has genomic data from population samples across the globe. You can enter the name of a country or population group, such as Siberian Yupik (the sample was collected from unrelated Siberian Yupiks from northeastern Siberia, Russia) and it will provide information on the number of people (29) and/or chromosomes sampled (58).

The criminal justice system began using DNA to solve crimes in the 1980s, after a geneticist from the University of Leicester in the UK developed a method for sequencing certain segments of chromosomal DNA. Those segments, called short tandem repeats (STRs), were expressed differently in different people, in contrast to the 99.9 percent of our DNA that is the same, and thus could be used to establish an indicator of personal identity (see Chapter 1).

As a recreational activity, with no serious consequences at stake, it barely matters whether the results consumers receive from their DNA ancestry tests accurately represent the percentages of their ancestry from different geographical regions. Given that there are no international standards for such testing, unlike genetic disease tests, it is not surprising that the results from different ancestry testing companies vary. As noted in Chapter 4, there are several stages in the analysis of a person’s saliva or cheek swabs where the criteria, reference frames, or analytics can vary among companies, yielding different outcomes.

When you purchase a DNA ancestry service you are sent a kit containing instructions for submitting a DNA sample. Most companies provide a plastic tube, which they ask the test-taker to fill with saliva or cheek swabs, seal, and return. When the company receives your DNA test sample it is processed for analysis. As noted previously, the vast amount of your genome does not distinguish you from other individuals. Therefore, your genome is broken down into segments of DNA that contain the alleles of interest, rather than it being fully sequenced. Here is how AncestryDNA describes the processing of the DNA samples it receives from customers: