Encrinurids are common in Ordovician and Silurian strata but whether they survived into the Early Devonian is still controversial. This paper documents the encrinurid Batocara sp. near the Silurian–Devonian boundary in western Junggar, Xinjiang. The highest horizon of Batocara sp. is located above the first appearance datum of the Devonian conodont Caudicriodus, confirming that encrinurids may cross the Silurian–Devonian boundary. The presence of Caudicriodus angustoides bidentatus, Zieglerodina planilingu and plate-type loboliths of scyphocrinoids above the highest horizon of Batocara sp. indicates that encrinurids here extend only into the lower part of the first conodont zone of the Lochkovian (i.e., Caudicriodus hesperius Biozone). Encrinurids are widely distributed and easily recognized, and unlike graptolites and conodonts are not controlled by lithofacies. Therefore, it might be possible to use the highest horizon of encrinurids as indicator fossils to identify the approximate position of the Silurian–Devonian boundary in areas or sections where graptolites and conodonts are not present, and at least in northwest China.

Late Kungurian (Lower Permian) conodonts are described from the Kanthan Limestone, Perak, Peninsular Malaysia and for the first time from the Sibumasu Terrane of the Malay Peninsula. The co-occurrence of Gullodus duani, Gullodus hemicircularis, Gullodus sicilianus, Mesogondolella lamberti and Mesogondolella siciliensis represent the Mesogondolella lamberti International Conodont Zone and the broadly equivalent Mesogondolella siciliensis Regional Conodont Zone. A small fault-bounded basal Pennsylvanian (basal Bashkirian) conodont fauna including Gnathodus girtyi simplex and Declinognathodus inaequalis is also reported. The late Kungurian conodonts from the Kanthan Limestone were deposited in a relatively deep-water environment on the northern passive margin of the Sibumasu Terrane of the eastern Cimmerian Continent located at c. 35oS latitude. Biogeographically, the fauna represents the southern peri-Gondwana Cool Water Province which is consistent with its palaeogeographic location. A new scheme, utilizing characteristics of P1 elements, including position of the 1st denticle, location of 2nd and 3rd denticles, platform shape, platform cross-section, denticle shape in cross-section, and lateral denticle development is proposed for distinguishing between species of the hindeodid genera Gullodus, Hindeodus and Isarcicella. The late Kungurian fauna from the Kanthan Limestone represents the southern peri-Gondwana Cool Water Province supporting palaeogeographic reconstructions placing the Sibumasu Terrane in moderate southern palaeolatitudes in the Kungurian.

The Viséan–Serpukhovian boundary is poorly defined in South China, hampering regional and global stratigraphical correlations. The foraminiferal and conodont distribution of the Baping Formation in the carbonate-slope Danlu section permits the recognition of an interval from the middle Viséan to the uppermost Serpukhovian in a continuous succession. The base of the Serpukhovian in Danlu is recognized by the first occurrences of Janischewskina delicata, Howchinia subplana and questionable ‘Millerella’ tortula. At a slightly younger level, the conodont Lochriea ziegleri is first recorded. A calibration on the first occurrence of L. ziegleri in different basins at a global scale has been revised compared to auxiliary markers within the ammonoids and foraminifers. The late occurrence of L. ziegleri in the Danlu section also supports a lack of synchronicity in the global first occurrence of this taxon. This study calls for the recognition of a new base for the Serpukhovian under a far better correlation between different zonal schemes and fossil groups.

Conodonts are an important biostratigraphic tool for many Phanerozoic stages. Along with graptolites, they define all global Ordovician Stage boundaries. Within the Upper Ordovician interval, a known species of Amorphognathus tvaerensis (Bergström) is present in both Sandbian and Katian stratotype sections. Study of changes in the succession of A. tvaerensis revealed that elements in the upper part of its range differ morphologically quite distinctly from those in its lower part. Here, they are described as a new conodont species, A. viirae sp. nov. This new species is recognized in several Estonian and Swedish sections, with apparent occurrence also in Mójcza Quarry, Holy Cross Mountains, Poland and Black Knob Ridge, Oklahoma, USA. Detailed analysis of early Amorphognathus elements from Estonian and Swedish sections revealed the absence of A. inaequalis (Rhodes) in both regions, although a conodont subzone based on this species was identified earlier by some authors. Both the absence of A. inaequalis (Rhodes) and recognition of the new species A. viirae sp. nov. resulted in the revision of the conodont zonation, and a new version of it is proposed for the Sandbian Stage in the Atlantic Realm. The new zonation includes (from below) Pygodus anserinus, Baltoniodus variabilis, A. tvaerensis, B. gerdae, A. viirae and B. alobatus Conodont zones.

Bennett Island stands alone in a remote part of the Arctic and information on its geology is essential to ascertain relations with other terranes in order to restore the early Palaeozoic Arctic palaeogeography. Lower Palaeozoic sedimentary rocks throughout the island were studied thoroughly for the first time. The Ordovician section (> 1.1 km thick) comprises three units: Tremadocian, lowest Floian black shale (130–140 m); Floian, lower Dapingian carbonate turbidite (> 250 m); and Dapingian, lower Darriwilian siliciclastic turbidite (> 730 m). Ordovician deposits conformably overlie Cambrian rocks deposited within the Siberian shelf, as shown earlier. Most of the Ordovician succession was formed in a deep trough that received carbonate debris from a nearby carbonate platform and silicate material from a distant landmass located to the NE (present coordinates). The Bennett Island Ordovician rocks have much in common with those of both the Central and Northern Taimyr belts. It could be tentatively suggested that both belts merged at their eastern continuation in the vicinity of De Long Islands. The whole system probably extends further eastwards. The Ordovician facies patterns and faunal assemblages in the New Siberian Islands are notably similar to those of northwestern Alaska, where the same lateral transition from turbidites to shelf limestones was reported.

The presence of a carbonate platform that interfingers towards the west with slope facies allows for the identification of an ancient lower Palaeozoic continental margin in the Western Precordillera of Argentina. The Los Sombreros Formation is essential for the interpretation of the continental slope of the Precordillera, which accreted to Gondwana as part of the Cuyania Terrane in the early Palaeozoic. The age of these slope deposits is controversial; therefore, a precise biostratigraphic scheme is critical to reveal the evolution of the South American continental margin of Gondwana. The study of lithic deposits of two sections of the Los Sombreros Formation, the El Salto and Los Túneles sections, provides important information for further understanding the depositional history of the slope. At El Salto section, the conodonts recovered from an allochthonous block refer to the Cordylodus proavus Zone (upper Furongian). The conodonts recovered from the matrix of a calclithite bed of the Los Sombreros Formation in the Los Túneles section are assigned to the Lenodus variabilis Zone (early Darriwilian), providing a minimum age for this stratigraphic unit. In addition, clasts from this sample yielded conodonts from the Paltodus deltifer − Macerodus dianae zones (upper Tremadocian). The contrasting conodont colour alterations and preservation states from the elements of two latter records, coming from the same sample, argue the reworked clasts originated in the carbonate platform and later transported to the slope during the accretion process of the Precordilleran Terrane to the South American Gondwanan margin during the Middle–Late Ordovician.

The Middle Shale Member of the Amdeh Formation is interpreted to be of Early Ordovician age based on its trace fossils, stratigraphic context and a newly discovered fauna of conodonts. The member abruptly overlies the Lower Quartzite Member, which may be Early Cambrian, and passes gradationally-upward into the Upper Quartzite Member, which is probably Early–Middle Ordovician. The 542.5 m thick Middle Shale Member can be divided into two parts: a shaly lower part, and a sandy upper part that contains an influx of heavy minerals. Bioturbation by marine trace fossils is one of the most obvious characteristics of the member. The shales and sandstones are interpreted to be of Cruziana and Skolithos ichnofacies and represent shallow-marine shelf, shoreface, beach and coastal deposits. Sparse shelly fossils occur in the sandy upper part, principally bivalves, inarticulate brachiopods, ostracods and conodonts. The small assemblage of conodonts includes elements interpreted to be Tremadocian (Tetraprioniodus, Drepanoistodus, Drepanodus, Scolopodus, ?Tropodus, Semiacontiodus and Teridontus), and others which may be Floian or ancestral forms of Floian taxa (Balognathidae gen. et sp. indet. A & B and aff. Erraticodon). No acritarchs have been recovered, probably due to high temperatures experienced during burial to >6 km. It is likely that the Middle Shale Member is the seaward equivalent of the Mabrouk and Barakat formations, and an outcrop gamma-ray log supports such a correlation. The trace fossils, sedimentology, conodont fauna and the general lack of macrofossils are in keeping with the regional Tremadocian–Floian of the Arabian margin of Gondwana.

Latest Sandbian to early Katian sequences across Laurentia's epicontinental sea exhibit a transition from lithologies characterized as ‘warm-water’ carbonates to those characterized as ‘cool-water'carbonates. This shift occurs across the regionally recognized M4/M5 sequence stratigraphic boundary and has been attributed to climatic cooling and glaciation, basin reorganization and upwelling of open ocean water, and/or increased water turbidity and terrigenous input associated with the Taconic tectophase. Documentation of oxygen isotopic trends across the M4/M5 and through bracketing strata provides a potential means of distinguishing among these alternative scenarios; however, oxygen isotopic records generated to date have failed to settle the debate. This lack of resolution is because δ18O records are open to multiple interpretations and potentially confounding factors related to local environmental conditions have not been tested by examining the critical interval in multiple areas and different depositional settings. To begin to address this shortcoming, we present new species-specific and mixed assemblage conodont δ18O values in samples spanning the M4/M5 boundary from the Upper Mississippi Valley, Alabama, and Virginia. The new results are combined with previous studies, providing a record of δ18O variability across SE Laurentia. The combined dataset allows us to test for regional trends at a resolution not previously available. Our results document a ~1.5‰ decrease in values across Laurentia instead of increasing δ18O values across the M4/M5 as predicted in various ‘cool-water’ scenarios. In short, these results do not support a shift to ‘cool-water’ conditions as an explanation for changes in early Katian carbonates across the M4/M5.

We have analysed a newly exposed conodont association from the Santa Gertrudis Formation in the Mojotoro Range, Eastern Cordillera of Salta province, Argentina. The key species Baltoniodus triangularis (Lindström), Trapezognathus quadrangulum (Lindström) and Erraticodon patu Cooper are present. A new species of the genus Baltoniodus (Baltoniodus cooperi sp. nov.) and elements of Periodon aff. P. flabellum are included in this conodont study. The key conodont B. triangularis indicates an age of lower Dapingian (Middle Ordovician) for the bearing strata. This was also recognized in other localities of the Eastern Cordillera, allowing their correlation. The conodonts from the Santa Gertrudis Formation provide conclusive elements on palaeogeographic interpretation by sharing species with Baltoscandia, South China and Australia, suggesting strong faunal affinities with these provinces.

The widely quarried and commercially exploited Gonioclymenia Limestone of the western and southern Tafilalt Platform (Anti-Atlas, SE Morocco) represents a transgressive but strongly condensed and shallowing upwards unit that overlies conformably or unconformably Famennian to Lower Devonian deposits. It is characterized by abundant, giant-sized Gonioclymenia speciosa, which characterizes the Famennian V-B, and rare Leviclymenia ramula sp. nov. Associated conodont faunas fall in the Bispathodus costatus Subzone of the B. aculeatus aculeatus (= Middle expansa) Zone. A rare outcrop at Jebel Ihrs West proved that Kalloclymenia and conodont faunas of the B. ultimus ultimus (= Upper expansa) Zone occur in a separate, overlying unit of the lower part of Famennian VI. Consequently, joint occurrences of Gonioclymenia and Kalloclymenia on large limestone slabs that are on offer in rock shops are artificial assemblages; there is no evidence for a regional co-occurrence of both genera. This is supported by correlation into thicker sections of the adjacent Maider and Tafilalt Basins. The Tafilalt Gonioclymenia Transgression may represent the regional expression of a eustatic pulse in the B. costatus Subzone that is known on the Ardennes Shelf as the transgressive Epinette Event. The conodont faunas yielded Neopolygnathus fibula sp. nov., Pseudopolygnathus primus tafilensis ssp. nov., new morphotypes of B. spinulicostatus and Pseudopolygnathus primus primus, the oldest Siphonodella (Eosiphonodella) and variable ‘siphonodelloids’ that are currently left in open nomenclature.

The Amdeh Formation is a 3.4 km stack of sparsely fossiliferous quartzites and shales which crops out in the Al Hajar mountains near Muscat. Here we describe the uppermost member (Am5) that can be dated biostratigraphically as Darriwilian and which is the outcrop equivalent, and probably the seaward continuation, of the Saih Nihayda Formation in the Ghaba Salt Basin of northern Oman. The outcrops at Wadi Daiqa and Hayl al Quwasim consist of 690 m of quartzitic sandstones, shales and bivalve-rich shell beds. Trace fossils referable to the Cruziana and Skolithos ichnofacies abound. The member comprises storm-dominated shelf, shoreface and delta deposits. A number of new discoveries have been made in the outcrops: fragments of the arandaspid fish Sacabambaspis, ossicles and moulds of the early disparid crinoid Iocrinus, two new genera of conodont, an occurrence of the rare trinucleid trilobite Yinpanolithus, and palynological and sedimentological evidence of more continuous Floian–Darriwilian deposition than is usual in the region. Sea levels during Middle Ordovician time are estimated to have been 50–200 m above present levels and a wide, low-gradient shelf covered much of Arabia. Similar trace fossils and storm-dominated, micro-tidal, sedimentary rocks occur throughout the region. Small changes of sea level, possibly caused by the growth and melting of polar ice sheets, could lead to substantial seaward or landward shifts of facies belts. The Am5 deposits are thick compared to most equivalents in Arabia implying active subsidence and a ready supply of sediment.

This contribution deals with the record of the Ordovician genus Pygodus in Cuyania, Argentina. Several classical sections have been sampled for conodonts in the Precordillera and the San Rafael Block, involving diverse sedimentary deposits: coarse clastic rocks with carbonate cement, carbonate beds intercalated in fine clastic deposits, green shale, and black carbonate deposits. The recovered species of this genus are Pygodus lunnensis Zhang, P. anitae Bergström, P. serra (Hadding), P. protoanserinus Zhang and P. anserinus Lamont & Lindström. These key conodonts indicate middle to upper Darriwilian – lower Sandbian age for the bearing strata. The species P. protoanserinus is recorded for the first time from the Precordillera. Detailed observations were made on specimens of P. anitae, P. lunnensis and P. protoanserinus; the two last species are described for the first time from Cuyania.

This paper is a contribution to the redefinition of the base of Carboniferous system. At present the criterion for the definition of the Devonian–Carboniferous boundary is the first occurrence of a conodont species. In order to evaluate the stratigraphic potential for new criteria for the definition of the Devonian–Carboniferous boundary, the distribution of conodont species of Bispathodus, Branmehla, Palmatolepis, Polygnathus, Protognathodus, Pseudopolygnathus and Siphonodella across the boundary is presented and discussed. An updated biozonation scheme across the boundary based on the First Appearance of Bispathodus ac. aculeatus, Bispathodus costatus, Bispathodus ultimus, Protognathodus kockeli, Siphonodella bransoni and Siphonodella duplicata is proposed, and it is suggested that the new criterion for the definition of the base of the Carboniferous system be the First Appearance Datum of Pr. kockeli or Si. bransoni.

One of the few and most complete records of the MDICE (Middle Darriwilian Isotope Carbon Excursion) is herein documented from Baltoscandia. Based on a core section penetrating the condensed Lower–Middle Ordovician succession (~46 m) on the island of Öland, southeastern Sweden, we provide an integrated scheme for carbon isotope chemostratigraphy (313 samples) and conodont biostratigraphy (29 samples) for this period. The carbonate succession in the Tingskullen core records 12 conodont zones and 6 subzones, including the Oepikodus evae, Trapezognathus diprion, Baltoniodus triangularis, B. navis, B. norrlandicus, Lenodus antivariabilis, L. variabilis, Yangtzeplacognathus crassus, Eoplacognathus pseudoplanus (Microzarkodina hagetiana and Microzarkodina ozarkodella subzones), E. suecicus, Pygodus serra (E. foliaceus, E. reclinatus, E. robustus and E. lindstroemi subzones) and Pygodus anserinus zones in ascending order. The δ13Ccarb record reveals an apparently complete record of the MDICE, including a rising limb, a well-defined peak and a falling limb. The anomaly covers a thickness of c. 27 m in the core and spans the Eoplacognathus pseudoplanus, E. suecicus, Pygodus serra and P. anserinus conodont zones. Combined with the new, detailed conodont biostratigraphy, the MDICE in the Tingskullen core can be used for detailed correlation with successions from Baltica, North America, the Argentine Precordillera, South China and North China.

Two stratigraphic sections spanning the mid-Carboniferous boundary were examined in the Clare Shale Formation of the Shannon Basin in Western Ireland. Calcareous nodules intermittently occur within these generally non-calcareous organic-rich shales, and these have yielded Serpukhovian and Bashkirian conodont elements. The biostratigraphic range of the Irish material is illustrated here for the first time. Results show that the mid-Carboniferous and Arnsbergian–Chokierian boundaries are coincident at Ballybunion. Gnathodus girtyi is restricted to the lower part of the Serpukhovian Stage (Late Mississippian), while G. bilineatus bollandensis persists into much younger strata, close to the first occurrence of Declinognathodus noduliferus s.l. One element belonging to G. postbilineatus is also present. These findings support the argument that D. noduliferus s.l. developed from G. b. bollandensis, rather than from G. girtyi. The biostratigraphical tool for the identification of the mid-Carboniferous boundary globally, and the suitability of the section at Arrow Canyon, USA as a GSSP, may therefore need to be reassessed.

We conducted isotope–geochemical studies of the Upper Devonian of the Rudny Altai. Whole-rock carbon isotope analyses of the studied section represented by carbonate fore-reef facies reveal a negative excursion at the base of punctata Zone. When compared to conodont biofacies turnovers and the Upper Devonian events chart, the new data have demonstrated a synchronism between the negative excursion and the coeval decline in conodont biodiversity corresponding to the Middlesex anoxic event of the Frasnian Stage of the Upper Devonian.

A conodont-graptolite biostratigraphic study was carried out on the top strata of the San Juan, Las Chacritas and Las Aguaditas formations in the La Trampa Range, Precordillera of San Juan in western Argentina. Significant conodont records in the San Juan and Las Chacritas formations allow for the recognition of the Yangtzeplacognathus crassus, Eoplacognathus pseudoplanus (Microzarkodina hagetiana and M. ozarkodella subzones) and Eoplacognathus suecicus zones of Darriwilian age. Index species and co-occurrences of graptolites and conodonts were recorded in the Las Aguaditas Formation allowing the identification of the Nemagraptus gracilis and the Pygodus anserinus zones, which represent the Sandbian Stage. These data indicate a hiatus between the Las Chacritas and the Las Aguaditas formations, corresponding to the Pygodus serra Zone and the Pterograptus elegans and Hustedograptus teretiusculus zones (upper Darriwilian). A total of 7287 identifiable conodont elements were recorded from the study section. The species frequency registered for each zone shows that Periodon and Paroistodus are the most abundant taxa, which are indicative of open marine environments. The records of particular conodont taxa, such as Histiodella, Periodon, Microzarkodina, Eoplacognathus and Baltoniodus, allow a precise global correlation with other regions such as south-central China, Baltoscandia, North America, Great Britain, Southern Australia and New Zealand. The graptolite fauna identified here are recognized worldwide in equivalent strata in the Baltic region, Great Britain, North America, China, southern Australia and New Zealand. The presence of graptolites in the ribbon limestones of the Las Chacritas Formation is documented for the first time.

The basement of the Central Pontides, and by implication that of Crimea, consists of pre-Permian low-grade metaclastic rocks intruded by latest Permian – Early Carboniferous (305–290 Ma) granitoids. Further up in the stratigraphic sequence are Triassic limestones, which are now preserved as olistoliths in the deformed Upper Triassic turbidites. New conodont and foraminifera data indicate an Anisian to Carnian (Middle to Late Triassic) age for these hemi-pelagic Hallstatt-type limestones. The siliciclastic turbidites surrounding the Triassic limestone contain the Norian (Late Triassic) bivalve Monotis salinaria; the same species is also found in the Tauric series in Crimea. The Upper Triassic flysch in the Central Pontides is locally underlain by basaltic pillow lavas and includes kilometre-size tectonic slices of serpentinite. Both the flysch and the serpentinite are cut by an undeformed acidic intrusion with an Ar–Ar biotite age of 162 ± 4 Ma (Callovian–Oxfordian). This indicates that the serpentinite was emplaced into the turbidites before Middle Jurassic time, most probably during latest Triassic or Early Jurassic time, and that the deformation of the Triassic sequence pre-dates the Middle Jurassic. Regional geological data from the circum-Black Sea region, including widespread Upper Triassic flysch, Upper Triassic eclogites and blueschists of oceanic crustal affinity, and apparent absence of a ‘Cimmerian continent’ between the Cretaceous and Triassic accretionary complexes indicate that the latest Triassic Cimmeride orogeny was accretionary rather than collisional and is probably related to the collision and accretion of an oceanic plateau to the southern active margin of Laurasia.

The selection of the global biomarker of the lower boundary of the Moscovian stages is one of the pressing issues of Carboniferous stratigraphy. Several solutions are suggested for this problem: Diplognathodus ellesmerensis Bender, Streptognathodus expansus (Igo & Koike) and Idiognathoides postsulcatus Nemirovskaya. The conodont species Declinognathodus donetzianus Nemirovskaya is one of the most prospective. It was detected in the rock sections of west Europe, the Donets Basin, the Moscow Syneclise, south Ural and the Appalachian Basin. The Volga region is also one of the places where Declinognathodus donetzianus Nemirovskaya is often met and this article is dedicated to detailed analysis of this species.

We present detailed biostratigraphy based on conodonts and palaeoenvironmental trends deduced from microfacies and conodont abundance through the Famennian (Late Devonian) at Col des Tribes (Montagne Noire, France). The succession is characterized by micritic limestones deposited in settings oscillating between mid to outer ramp. Facies contain poor fauna, widely dominated by nektonic organisms. This section is complete and one of the most conodont-rich for the Famennian of the north Gondwana-related area. The Upper Kellwasser event (Frasnian–Famennian boundary) and the Hangenberg (Devonian–Carboniferous boundary) have been lithologically identified. They are characterized by decimetre-thick black dysoxic to anoxic argillaceous sediments. The Condroz and annulata events, although not materialized by lithological changes, have been positioned due to the precise stratigraphy. The first event occurred during the deposition of condensed ferruginous facies (griotte limestones) and the second event during the deposition of micrites barren of benthic fauna. The combination of information from both facies and conodont biofacies changes allows a general sea-level curve through the entire Famennian for north Gondwana to be proposed for the first time. At Col des Tribes, the general trend is a slight deepening upwards from triangularis to trachytera zones, then a pronounced shallowing-upwards trend from upper trachytera to praesulcata zones. This curve correlates with the well-known reference curve from Euramerica concerning the late Famennian (trachytera to praesulcata Zones). There are some discrepancies in minor cycles which can be explained by tectonical phenomena at the onset of the edification of the Variscan belt in Europe.