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The Early Ordovician Middle Shale Member (Am3) of the Amdeh Formation and further evidence of conodont faunas from the Sultanate of Oman

Published online by Cambridge University Press:  26 November 2018

AP Heward*
Affiliation:
23 Croftdown Court, Malvern, WR14 3HZ, UK
CG Miller
Affiliation:
Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK
GA Booth
Affiliation:
Weyhill, Old Compton Lane, Farnham, Surrey GU9 8EG, UK
*
Author for correspondence: AP Heward, Email: alan@midfarm.demon.co.uk

Abstract

The Middle Shale Member of the Amdeh Formation is interpreted to be of Early Ordovician age based on its trace fossils, stratigraphic context and a newly discovered fauna of conodonts. The member abruptly overlies the Lower Quartzite Member, which may be Early Cambrian, and passes gradationally-upward into the Upper Quartzite Member, which is probably Early–Middle Ordovician. The 542.5 m thick Middle Shale Member can be divided into two parts: a shaly lower part, and a sandy upper part that contains an influx of heavy minerals. Bioturbation by marine trace fossils is one of the most obvious characteristics of the member. The shales and sandstones are interpreted to be of Cruziana and Skolithos ichnofacies and represent shallow-marine shelf, shoreface, beach and coastal deposits. Sparse shelly fossils occur in the sandy upper part, principally bivalves, inarticulate brachiopods, ostracods and conodonts. The small assemblage of conodonts includes elements interpreted to be Tremadocian (Tetraprioniodus, Drepanoistodus, Drepanodus, Scolopodus, ?Tropodus, Semiacontiodus and Teridontus), and others which may be Floian or ancestral forms of Floian taxa (Balognathidae gen. et sp. indet. A & B and aff. Erraticodon). No acritarchs have been recovered, probably due to high temperatures experienced during burial to >6 km. It is likely that the Middle Shale Member is the seaward equivalent of the Mabrouk and Barakat formations, and an outcrop gamma-ray log supports such a correlation. The trace fossils, sedimentology, conodont fauna and the general lack of macrofossils are in keeping with the regional Tremadocian–Floian of the Arabian margin of Gondwana.

Type
Original Article
Copyright
© Cambridge University Press 2018 

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References

Abdulkadir, IT and Abdullatif, OM (2013) Facies, depositional environments, reservoir potential and palaeogeography of the Cambro-Ordovician Dibsiyah formation, Wajid outcrop belt, Saudi Arabia. Arabian Journal of Science and Engineering 38, 1785–806.CrossRefGoogle Scholar
Agematsu, S, Sashida, K, Salyapongse, S and Sarsud, A (2007) Ordovician conodonts from the Satun Area, Southern Peninsula Thailand. Journal of Paleontology 81, 1937.CrossRefGoogle Scholar
Al-Hadidy, AH (2007) Paleozoic stratigraphic lexicon and hydrocarbon habitat of Iraq. GeoArabia 12, 63130.Google Scholar
Aplin, AC and Macquaker, JHS (2011) Mudstone diversity: origin and implications for source seal and reservoir properties in petroleum systems. Bulletin of the American Association of Petroleum Geologists 95, 2031–59.CrossRefGoogle Scholar
Bagnoli, G, Machado, G and Al-Marjibi, S (2016) The first record of Cambrian conodonts from the Huqf-Haushi outcrops, Oman, Arabian Peninsula. Rivista Italiana di Paleontologia e Stratigrafia 122, 235–42.Google Scholar
Bagnoli, G and Stouge, S (1996) Lower Ordovician (Billingen-Kunda) conodont zonation and provinces based on sections from Horns Ude, north Öland, Sweden. Bollettino della Società Paleontologica Italiana 35, 109–63.Google Scholar
Bagnoli, G, Stouge, S and Tongiorgi, SM (1988) Acritarchs and conodonts from the Cambro-Ordovician Furuhäll (Köpings-klint) section (Öland, Sweden). Rivista Italiana di Paleontologica e Stratigrafia 94, 163248.Google Scholar
Bergström, SM and Ferretti, A (2016) Conodonts in Ordovician biostratigraphy. Lethaia 50, 424–39.CrossRefGoogle Scholar
Bottjer, DJ and Droser, ML (1991) Ichnofabric and basin analysis. Palaios 6, 199205.CrossRefGoogle Scholar
Breton, J-P, Béchennec, F, Le Métour, J, Moen-Maurel, L and Razin, P (2004) Eoalpine (Cretaceous) evolution of the Oman Tethyan continental margin: insights from a structural field study in Jabal Akhdar (Oman Mountains). GeoArabia 9, 4158.Google Scholar
Burkhard, M (1993) Calcite twins, their geometry, appearance and significance as stress-strain markers and indicators of tectonic regime: a review. Journal of Structural Geology 15, 351–68.CrossRefGoogle Scholar
Carlorosi, JMT (2013) La Zona de Baltoniodus triangularis (Conodonta) en el Paleozoico de la Cuenca Central Andina Sudamericana: Formación Alto del Cóndor del Norte Argentino. Boletín Geológico y Minero 124, 551–62.Google Scholar
Carlorosi, JMT and Heredia, SE (2013) The Ordovician conodont Trapezognathus Lindström, 1955 in the Andean Basin, Argentina. Neues Jahrbuch für Geologie und Paläontologie – Abhandlungen 267, 309–21.CrossRefGoogle Scholar
Carlorosi, J, Heredia, S and Aceñolaza, G (2013) Middle Ordovician (early Dapingian) conodonts in the central Andean Basin of NW Argentina. Alcheringa: An Australasian Journal of Palaeontology 37, 299311.CrossRefGoogle Scholar
Chauvet, F, Dumont, T and Basile, C (2009) Structures and timing of Permian rifting in the central Oman Mountains (Saih Hatat). Tectonophysics 475, 563–74.CrossRefGoogle Scholar
Clifton, HE (1969) Beach lamination: nature and origin. Marine Geology 7, 553–9.CrossRefGoogle Scholar
Cornish, S and Searle, MP (2017) 3D geometry and kinematic evolution of the Wadi Mayh sheath fold, Oman, using detailed mapping from high-resolution photography. Journal of Structural Geology 101, 2642.CrossRefGoogle Scholar
Dean, WT (2004) Cambrian and Ordovician correlation and trilobite distribution in Turkey. Fossils and Strata 5, 353–73.Google Scholar
Droste, HHJ (1997) Stratigraphy of the lower Paleozoic Haima Supergroup of Oman. GeoArabia 2, 419–72.Google Scholar
Dzik, J (2015) Evolutionary roots of the conodonts with increased number of elements in the apparatus. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 106, 2953.CrossRefGoogle Scholar
Epstein, A, Epstein, JB and Harris, LD (1977) Conodont color alteration – an index to thermal metamorphism. Geological Society of America Professional Paper 95, 27 pp.CrossRefGoogle Scholar
Forbes, GA, Jansen, HSM and Schreurs, J (2010) Haima Supergroup. In Lexicon of Oman Subsurface Stratigraphy, pp. 171203. Manama, Bahrain: GeoArabia, Special Publication no. 5.Google Scholar
Fortey, RA (1994) Late Cambrian trilobites from the Sultanate of Oman. Neues Jahrbuch für Geologie und Paläontologie – Abhandlungen 194, 2553.Google Scholar
Fortey, RA, Heward, AP and Miller, CG (2011) Sedimentary facies and trilobite and conodont faunas of the Ordovician Rann formation, Ras al Khaimah, United Arab Emirates. GeoArabia 16, 127–52.Google Scholar
Fortey, RA and Seilacher, A (1997) The trace fossil Cruziana semiplicata and the trilobite that made it. Lethaia 30, 105–12.CrossRefGoogle Scholar
Fraser, WT, Watson, JS, Sephton, MA, Lomax, BH, Harrington, G, Gosling, WD and Self, S (2014) Changes in spore chemistry and appearance with increasing maturity. Review of Palaeobotany and Palynology 201, 41–6.CrossRefGoogle Scholar
Ghavidel-Syooki, M, Popov, LE, Álvaro, JJ, Ghobadi, Pour M, Tolmacheva, TY and Ehsani, M-H (2014) Dapingian–lower Darriwilian (Ordovician) stratigraphic gap in the Faraghan mountains, Zagros ranges, south-eastern Iran. Bulletin of Geosciences, Czech Geological Survey 89, 679706.Google Scholar
Ghavidel-Syooki, M and Vecoli, M (2008) Palynostratigraphy of middle Cambrian to lowermost Ordovician stratal sequence in the High Zagros mountains, southern Iran: regional stratigraphic implications and palaeobiogeographic significance. Review of Palaeobotany and Palynology 150, 97114.CrossRefGoogle Scholar
Ghobadi, Pour M (2006) Early Ordovician (Tremadocian) trilobites from Simeh-Kuh, Eastern Alborz, Iran. In Studies in Palaeozoic Palaeontology (eds Bassett, MG and Deisler, VK), pp. 93118. Cardiff: National Museum of Wales Geological Series no. 25.Google Scholar
Ghobadi, Pour M, Kebriaee-Zadeh, MR and Popov, LE (2011a) Early Ordovician (Tremadocian) brachiopods from the Eastern Alborz mountains, Iran. Estonian Journal of Earth Sciences 60, 6582.Google Scholar
Ghobadi, Pour M, Mohibullah, M, Williams, M, Popov, LE and Tolmacheva, TY (2011b) New early ostracods from the Ordovician (Tremadocian) brachiopods of Iran: systematic, biogeographical and palaeoecological significance. Alcheringa 35, 517–29.CrossRefGoogle Scholar
Hansman, RJ, Ring, U, Thompson, SN, den Brok, B and Stübner, K (2017) Late Eocene uplift of the Al Hajar mountains, Oman, supported by stratigraphy and low-temperature thermochronology. Tectonics 36, 3081–109.CrossRefGoogle Scholar
Haq, BU and Schutter, SR (2008) A chronology of Paleozoic sea-level changes. Science 322, 64–8.CrossRefGoogle ScholarPubMed
Heredia, S, Carlorosi, J, Mestre, A and Soria, T (2013) Stratigraphical distribution of the Ordovician conodont Erraticodon Dzik in Argentina. Journal of South American Earth Sciences 45, 224–34.CrossRefGoogle Scholar
Heward, AP and Penney, RA (2014) Al Khlata glacial deposits in the Oman Mountains and their implications. In Tectonic Evolution of the Oman Mountains (eds. Rollinson, HR, Searle, MP, Abbasi, IA, Al-Lazki, A and Al Kindi, MH), pp. 279301. Geological Society of London, Special Publication no. 392.Google Scholar
Heward, AP, Booth, GA, Fortey, RA, Miller, CG and Sansom, IJ (2018) Darriwilian shallow-marine deposits from the Sultanate of Oman, a poorly known portion of the Arabian margin of Gondwana. Geological Magazine 155, 5984.CrossRefGoogle Scholar
Horowitz, AS and Potter, PE (1971) Introductory Petrography of Fossils. New York: Springer-Verlag, 302 pp.CrossRefGoogle Scholar
Hughes, Clarke MW (1988) Stratigraphy and rock unit nomenclature in the oil-producing area of interior Oman. Journal of Petroleum Geology 11, 560.CrossRefGoogle Scholar
Jahangir, H, Ghobadi, Pour M, Ashuri, A-R and Amini, A (2015) Terminal Cambrian and early Ordovician (Tremadocian) conodonts from Eastern Alborz, north-central Iran. Alcheringa 40, 219–43.CrossRefGoogle Scholar
Jahangir, H, Ghobadi, Pour M, Holmer, LE, Popov, LE, Ashuri, A-R, Rushton, A, Tolmacheva, TY and Amini, A (2014) Biostratigraphy of the Cambrian-Ordovician boundary beds at Kopet-Dagh, Iran. Stratigraphy 12, 40–7.Google Scholar
Jahangir, H, Ghobadi Pour, M, Tolmacheva, TY, Popov, L and Hosseini-Nezhad, M (2012) Conodont and trilobite biostratigraphy across the Cambrian-Ordovician boundary in Deh-Molla eastern Alborz, Iran. Palaeontological Association Newsletter 81, 70–1.Google Scholar
Ji, Z and Barnes, CR (1996) Uppermost Cambrian and lower Ordovician conodont biostratigraphy of the survey peak formation (Ibexian/Tremadoc), Wilcox Pass, Alberta, Canada. Journal of Paleontology 70, 871–90.CrossRefGoogle Scholar
Kidwell, SM, Fursich, FT and Aigner, T (1986) Conceptual framework for the analysis and classification of fossil concentrations. Palaios 1, 228–38.CrossRefGoogle Scholar
Knox, RWO’B, Franks, SG and Cocker, JD (2007) Stratigraphic evolution of heavy-mineral provenance signatures in the sandstones of the Wajid group (Cambrian to Permian), southwestern Saudi Arabia. GeoArabia 12, 6596.Google Scholar
Konert, G, Afifi, AA, Al-Hajri, SA and Droste, HJ (2001) Paleozoic stratigraphy and hydrocarbon habitat of the Arabian plate. GeoArabia 6, 407–42.Google Scholar
Lehnert, O (1995) Ordovizische Conodonten aus der Präkordillere Westargentiniens: Ihre Bedeutung für Stratigraphie und Paläogeographie. Erlanger Geologische Abhandlung 125, 1193.Google Scholar
Lehnert, O, Nowak, H, Sarmiento, GN, Gutiérrez-Marco, JC, Akodad, M and Servais, T (2016) Conodonts from the lower Ordovician of Morocco – contributions to age and faunal diversity of the Fezouata Lagerstätte and peri-Gondwana biogeography. Palaeogeography, Palaeoclimatology, Palaeoecology 460, 5061.CrossRefGoogle Scholar
Le Métour, J, Villey, M and De Gramont, X (1986) Geological Map of Qurayat, Sheet NF 40-4D, Scale 1:100,000. Explanatory Notes. Oman: Directorate General of Minerals, Oman Ministry of Petroleum and Minerals, 72 pp.Google Scholar
Lindström, M (1955) Conodonts from the lowermost Ordovician strata of south-central Sweden. Geologiska Föreningens i Stockholm Förhandlingar 76, 517604.CrossRefGoogle Scholar
Löfgren, A (1978) Arenigian and Llanvirnian conodonts from Jämtland, northern Sweden. Fossils and Strata 13, 129.Google Scholar
Löfgren, A (1993) Conodonts from the lower Ordovician at Hunneberg, south-central Sweden. Geological Magazine 130, 215–32.CrossRefGoogle Scholar
Löfgren, A (1994) Arenig (lower Ordovician) conodonts and biozonation in the eastern Siljan district, central Sweden. Journal of Paleontology 68, 1350–68.CrossRefGoogle Scholar
Löfgren, A (1999) The Ordovician conodont Semiacontiodus cornuformis (Sergeeva, 1963). Geologica et Palaeontologica 33, 7191.Google Scholar
Lovelock, PER, Potter, TL, Walsworth-Bell, EB and Wiemer, WM (1981) Ordovician rocks in the Oman mountains: the Amdeh formation. Geologie en Mijnbouw 60, 487–95. (Also 1:1000 Composite Section, Amdeh Formation, Wadi Kahza – Wadi Amdeh, Saih Hatat, 1979, unpublished).Google Scholar
Mattern, F, Pracejus, B and Al-Balushi, L (2018). Heavy mineral beach placers of the Ordovician Amdeh formation (Member 4, Wadi Qahza, Saih Hatat, eastern Oman Mountains): where is the main source area? Journal of African Earth Sciences 147, 633–46.CrossRefGoogle Scholar
McCracken, A and Nowlan, G (1989) Conodont paleontology and biostratigraphy of Ordovician and petroliferous carbonates from Southampton, Baffin and Aptapok islands in the eastern Canadian Arctic. Canadian Journal of Earth Sciences 26, 1880–903.CrossRefGoogle Scholar
Miller, CG, Heward, AP, Mossoni, A and Sansom, IJ (2018). Two new early balognathid conodont genera from the Ordovician of Oman and comments on the early evolution of prioniodontid conodonts. Journal of Systematic Palaeontology 16, 571–93.CrossRefGoogle Scholar
Millson, JA, Quin, JG, Idiz, E, Turner, P and Al-Harthy, A (2008) The Khazzan gas accumulation, a giant combination trap in the Cambrian Barik Sandstone member, Sultanate of Oman: implications for Cambrian petroleum systems and reservoirs. Bulletin of the American Association of Petroleum Geologists 92, 885917.CrossRefGoogle Scholar
Molyneux, SG and Al-Hajri, S (2000) Palynology of a problematic Lower Palaeozoic lithofacies in central Saudi Arabia. In Stratigraphic Palynology of the Palaeozoic of Saudi Arabia (eds Al-Hajri, S and Owens, B), pp. 1841. Manama, Bahrain: GeoArabia Special Publication no. 1.Google Scholar
Molyneux, SG, Osterloff, P, Penney, RA and Spaak, P (2006) Biostratigraphy of the lower Palaeozoic Haima supergroup, Oman; its application in sequence stratigraphy and hydrocarbon exploration. GeoArabia 11, 1748.Google Scholar
Mount, VS, Crawford, RIS and Bergman, SC (1998) Regional structural style of the central and southern Oman mountains: Jebel Akhdar, Saih Hatat and the northern Ghaba Salt basin. GeoArabia 3, 475–90.Google Scholar
Morton, DM (1959) The geology of Oman. Proceedings of the 5th World Petroleum Congress, vol. 1, pp. 277–94. New York.Google Scholar
Polechová, M (2016) The bivalve fauna from the Fezouata formation (lower Ordovician) of Morocco and its significance for palaeobiogeography, palaeoecology and early diversification of bivalves. Palaeogeography, Palaeoclimatology, Palaeoecology 460, 155–69.CrossRefGoogle Scholar
Popov, LE, Ghobadi, Pour M, Bassett, MG and Kebriaee-Zadeh, MR (2009) Billingsellide and orthide brachiopods: new insights into earliest Ordovician evolution and biogeography from Northern Iran. Palaeontology 52, 3552.CrossRefGoogle Scholar
Rasmussen, JA (2001) Conodont biostratigraphy and taxonomy of the Ordovician shield margin in the Scandinavian Caledonides. Fossils and Strata 48, 1180.Google Scholar
Rickards, BR, Booth, GA, Paris, F and Heward, AP (2010) Marine flooding events of the early and middle Ordovician of Oman and the United Arab Emirates and their graptolite, acritarch and chitinozoan associations. GeoArabia 15, 81120.Google Scholar
Rickards, BR, Hamedi, MA and Wright, AJ (1994) A new Arenig graptolite fauna from the Kerman district, east-central Iran. Geological Magazine 131, 3542.CrossRefGoogle Scholar
Roy, PS (1999) Heavy mineral beach placers in southeastern Australia; their nature and genesis. Economic Geology 94, 567–88.CrossRefGoogle Scholar
Saddiqi, O, Michard, A, Goffe, B, Poupeau, G and Oberhänsli, R (2006) Fission-track thermochronology of the Oman mountains continental windows, and current problems of tectonic interpretation. Bulletin de la Société géologique de France 177, 127–34.CrossRefGoogle Scholar
Searle, MP, Warren, CJ, Waters, DJ and Parish, RR (2004) Structural evolution of the Arabian continental margin, Saih Hatat region, Oman mountains. Journal of Structural Geology 26, 451–73.CrossRefGoogle Scholar
Seilacher, A (1962) Form und Function des Trilobiten-daktylus. Paläontologische Zeitschrift 36, 218–27.CrossRefGoogle Scholar
Seilacher, A (1970) Cruziana stratigraphy of ‘non-fossiliferous’ Palaeozoic sandstones. In Trace Fossils (eds Crimes, TP and Harper, JC), pp. 447–76. Geological Journal Special Issue no. 3. Liverpool: Seel House Press.Google Scholar
Seilacher, A (1991) An updated Cruziana stratigraphy of Gondwanan Palaeozoic sandstones. In The Geology of Libya IV (eds Salem, MJ, Hammuda, OS and Eliagoubi, BA), pp. 1565–81. Amsterdam: Elsevier.Google Scholar
Seilacher, A (2000) Ordovician and Silurian arthrophycid ichnostratigraphy. In Geological Exploration in Murzuq Basin (eds Sola, MA and Worsley, D), pp. 237–58. Amsterdam: Elsevier Science B.V.CrossRefGoogle Scholar
Seilacher, A (2007) Trace Fossil Analysis. Berlin; Heidelberg: Springer-Verlag, 226 pp.Google Scholar
Seilacher, A (2008) Fossil Art. Laasby: CBM-Publishing, 102 pp.Google Scholar
Selley, RC (1970) Ichnology of Palaeozoic sandstones in the southern Desert of Jordan: a study of trace fossils in their sedimentological context. In Trace Fossils (eds Crimes, TP and Harper, JC), pp. 477–88. Geological Journal Special Issue no. 3. Liverpool: Seel House Press.Google Scholar
Selley, RC (1972) Diagnosis of marine and non-marine environments from the Cambro-Ordovician sandstones of Jordan. Journal of the Geological Society 128, 135–50.CrossRefGoogle Scholar
Serpagli, E (1974) Lower Ordovician conodonts from precordilleran Argentina (province of San Juan). Bollettino della Società Paleontologica Italiana 13, 1798.Google Scholar
Serpagli, E, Ferretti, A, Nicoll, RS and Serventi, P (2008) The conodont genus Teridontus (Miller, 1980) from the early Ordovician of the Montagne Noire, France. Journal of Paleontology 82, 612–20.CrossRefGoogle Scholar
Stouge, S and Bagnoli, G (1988) Early Ordovician conodonts from the Cow Head Peninsula, western Newfoundland. Palaeontographia Italica 75, 89179.Google Scholar
Stouge, S and Bagnoli, G (1990) Lower Ordovician (Volkhovian-Kundan) conodonts from Hagudden, northern Öland, Sweden. Palaeontographia Italica 77, 154.Google Scholar
Stouge, S and Bagnoli, G (1999) The suprageneric classification of some Ordovician prioniodontid conodonts. Bollettino della Società Paleontologica Italiana 37, 145–58.Google Scholar
Tolmacheva, TY (2006) Apparatus of the conodont Scolopodus striatus Pander, 1856 and a re–evaluation of Pander’s species of Scolopodus. Acta Palaeontologica Polonica 51, 247–60.Google Scholar
Turner, P (1979) Diagenetic origin of Cambrian marine red beds: Caerfai Bay Shales, Dyfed, Wales. Sedimentary Geology 24, 269–81.CrossRefGoogle Scholar
van Wamel, WA (1974) Conodont biostratigraphy of the upper Cambrian and lower Ordovician of north western Öland. Utrecht Micropalaeontological Bulletins 10, 126.Google Scholar
Vaslet, D (1990) Le Paléozoïque (anté-Permien supérieur) d’Arabie Saoudite. Document du BRGM no. 191, 209 pp.Google Scholar
Vaucher, R, Pittet, B, Hormière, H, Martin, LO and Lefebvre, B (2017) A wave-dominated, tide-modulated model for the lower Ordovician of the Anti-Atlas, Morocco. Sedimentology 64, 777807.CrossRefGoogle Scholar
Viira, V, Aldridge, RJ and Curtis, S (2006a) Conodonts of the Kiviõli member, Viivikonna formation (upper Ordovician) in the Kohtla section, Estonia. Proceedings of the Estonian Academy of Science, Geology 55, 213–40.Google Scholar
Viira, V, Mens, K and Nemliher, J (2006b) Lower Ordovician Leetse formation in the North Estonian Klint area. Proceedings of the Estonian Academy of Science, Geology 55, 156–74.Google Scholar
Viira, V, Löfgren, A, Mägi, S and Wickström, J (2001) An early to middle Ordovician succession of conodont faunas at Mäekalda, northern Estonia. Geological Magazine 138, 699718.CrossRefGoogle Scholar
Villey, M, Le Métour, J and De Gramont, X (1986) Geological Map of Fanjah, Sheet NF 40-3F, Scale 1:100,000. Explanatory Notes. Oman: Directorate General of Minerals, Oman Ministry of Petroleum and Minerals, 68 pp.Google Scholar
Voldman, GG, Albanesi, GL, Ortega, G, Giuliano, ME and Monaldi, CR (2017) New conodont taxa and biozones from the lower Ordovician of the Cordillera Oriental, NW Argentina. Geological Journal 52, 394414.CrossRefGoogle Scholar
Walker, RG and Plint, AG (1992) Wave and storm-dominated shallow marine systems. In Facies Models: Response to Sea Level Change (eds Walker, RG and James, NP), pp. 219–38. Toronto: Geological Association of Canada.Google Scholar
Weber, KJ (1971) Sedimentological aspects of the oil fields in the Niger delta. Geologie en Mijnbouw 50, 559–76.Google Scholar
Wellman, CH, Osterloff, PL and Mohiuddin, U (2003) Fragments of the earliest land plants. Nature 425, 282–5.CrossRefGoogle ScholarPubMed
Zhen, YY and Percival, IG (2003) Ordovician conodont biogeography – reconsidered. Lethaia 36, 357–70.CrossRefGoogle Scholar
Ziegler, AM and McKerrow, WS (1975) Silurian marine red beds. American Journal of Science 275, 3156.CrossRefGoogle Scholar