A study was conducted to evaluate effects of application timing and sulfometuron rate on weed control and bermudagrass tolerance. Sulfometuron was applied at rates from 13 to 105 g ai/ha and compared to 1120 g ae/ha 2,4-D amine. Control of Italian ryegrass and common vetch was not affected by October, November, December, or March application dates and 39 to 105 g/ha sulfometuron provided equivalent control of both species 6 mo after the initial treatment. Percent green cover of bermudagrass at 6 mo after the initial treatment was 22% or less following March applications compared to 36% or greater when applied in October, November, or December. More bermudagrass injury occurred following March treatments of 52 g/ha or greater sulfometuron where green cover was 14% and less compared to 22 and 30% green cover following 13 g/ha sulfometuron or 1120 g/ha 2,4-D amine, respectively.

Greenhouse experiments in central Texas assessed the relative importance of above- and belowground interactions of semidwarf Mit wheat and Marshall ryegrass during vegetative growth. One experiment used partitions to compare the effect of no (controls), aboveground only, belowground only, and full interaction for 75 d after planting (DAP) one wheat and nine ryegrass plants in soil volumes of 90, 950, and 3,800 ml. The results with the different soil volumes were similar. Wheat growth in the aboveground interaction only did not differ from controls. However, the full or belowground only interaction of wheat with ryegrass reduced wheat height, leaf number, tillering, leaf area, percent total nonstructural carbohydrates in shoot, and dry weights of leaves, stems, and roots 45 and 75 DAP compared to controls. Wheat in full and belowground interaction only did not differ from one another in growth. A replacement series experiment of 56 d also showed that the competitive advantage of ryegrass was relatively greater in root than in shoot growth. No allelopathic response of wheat to ryegrass occurred. While the tallness of the semidwarf wheat minimized aboveground interference by ryegrass, the root growth of the thinner and more fibrous roots of ryegrass greatly enhanced its belowground competitiveness.

Flower production and pollen dispersal patterns of diclofop-methyl-susceptible and -resistant Italian ryegrass were examined to determine whether evolution of herbicide resistance could be controlled by cross-fertilization between biotypes. The susceptible and resistant ryegrass biotypes differed in timing and abundance of both ovule production and pollen release such that pollen from the susceptible plants had a much greater chance of fertilizing the resistant plant population than vice versa. Susceptible ryegrass, growing with or without wheat competition, produced more than 60% of its seeds before any pollen from resistant plants was released. In contrast, throughout the course of resistant plant seed generation, pollen from susceptible plants composed at least 30% of the total pollen load. These phenological differences, in conjunction with herbicide treatment results for third-generation ryegrass seedlings, suggest that gene flow can be used to reduce development of diclofop-methyl resistance in ryegrass populations within wheat cropping systems. In a wheat field infested with resistant ryegrass, cessation of herbicide application and sowing of a susceptible ryegrass biotype can be expected to reduce the evolution of diclofop-methyl resistance by at least 6% per year.

The efficacy of herbicides applied broadcast by hand with a soil carrier or sprayed with water carrier were compared for Italian ryegrass control in winter wheat in central Oklahoma. Italian ryegrass was controlled 77 to 100% with BAY SMY 1500, chlorsulfuron, and CGA-131036 applied with both carriers. Metribuzin applied POST with 187 L ha–1 water was less phytotoxic to Italian ryegrass and wheat than with 200 kg ha–1 soil. All herbicides applied with 200 kg ha–1 silt loam soil as carrier at two locations controlled Italian ryegrass 96 to 100%. Increasing herbicide rate, soil carrier quantity, or broadcasting soil-carried herbicide in two directions rather than one did not significantly improve herbicide efficacy over that of conventional spraying.

Research was conducted to determine the potential for allelopathy to occur in pastures infested with bitter sneezeweed. Aqueous extracts of bitter sneezeweed leaves reduced alfalfa and Italian ryegrass seedling growth as much as 50% at concentrations of 0.5% (w/v). Leaf extracts were more phytotoxic than either stem or root extracts and seedling growth was reduced more than seed germination. Bitter sneezeweed tissue mixed in potting soil at concentrations as low as 0.3% w/w reduced alfalfa seedling numbers by 43%, plant height by 26%, and foliage dry matter production by 54% compared to plants cultured in soil without bitter sneezeweed leaf tissue. The potential concentration of bitter sneezeweed leaf material in soil in the pasture ecosystem was determined to be 0.5% w/v in the liquid phase and 0.2% w/w in the solid phase. Alfalfa seedling growth was reduced by 70% when germinating seed were placed under a bell jar with a potted mature bitter sneezeweed plant compared to control seedlings. A potential exists for bitter sneezeweed interference with developing alfalfa and Italian ryegrass seedlings following fall interseeding into pastures infested with bitter sneezeweed.

Above-ground seedling development was characterized for five annual grass weeds: downy brome, bulbous bluegrass, jointed goatgrass, Italian ryegrass, wild oat; and three cereals: winter wheat, winter barley, and winter triticale in field experiments over two years. The rate of leaf production on the main stem of each species was linearly related to cumulative growing degree days (GDD) since planting. Leaf production rates were faster for bulbous bluegrass, downy brome, Italian ryegrass, wild oat, and barley than for wheat, triticale, and jointed goatgrass. The main stem development stage when individual tillers appeared was similar in all species except under poor seedbed conditions in 1991, in which case lower-node tillers were delayed in the cereals and jointed goatgrass, but not in most of the weed species. Bulbous bluegrass, downy brome, and barley had the same percentage of plants produce the first four primary tillers on the main stem in both years; the other species showed more year-to-year variation. Seedling heights at full emergence were generally greater for large-seeded species. Small-seeded species compared to large-seeded species tended to have greater relative increases in plant height over time. Knowledge of comparative development rates between these weeds and cereals could provide information for development of growth models for each of the species and could also improve understanding of the competitive relationships between grass weeds and cereal grains.

Field experiments were conducted in 1982 through 1986 to evaluate the effect of six postemergence grass herbicides, clethodim, the methyl ester of diclofop, the butyl ester of fluazifop-P, the methyl ester of haloxyfop, the methyl ester of quizalofop, and sethoxydim, on 31 seedling grass species. All herbicides except diclofop controlled most of the species tested. None of the herbicides controlled rattail fescue or red fescue. Only clethodim and haloxyfop controlled annual bluegrass.

In the field, 14 soft red winter wheat cultivars responded differently to 1.1 kg ai ha–1 diclofop, 1.7 kg ai ha–1 BAY SMY 1500, and 0.42 kg ai ha–1 metribuzin applied POST. Diclofop and metribuzin did not injure any cultivar more than 10% on a silty clay soil. However, BAY SMY 1500 injured ‘Pioneer 2551’ and ‘Coker 983’ 39 and 21%, respectively, in March. All other cultivars were injured less than 10% by BAY SMY 1500. Early injury did not translate into yield loss in the cultivar tolerance study. In an application timing study for Italian ryegrass control, late-season ratings indicated better control with two-leaf applications than with PRE applications for all treatments. Delaying application to the three-tiller stage reduced control with BAY SMY 1500 or metribuzin, but not with diclofop. On the sandier soil at this location, wheat injury with 0.28 or 0.43 kg ha–1 metribuzin or 2.2 kg ha–1 BAY SMY 1500 was sufficient to reduce wheat yield compared with other treatments, despite good Italian ryegrass control.

Research was conducted on a sandy clay loam in Georgia during 1987–88 and 1988–89 to determine winter wheat, oats, barley, and Italian ryegrass tolerance to flurtamone. Flurtamone was applied PPI, PRE, and early POST at 0.28, 0.56, and 1.1 kg ai ha-1. Order of tolerance of the four species was barley > wheat > oat > Italian ryegrass. Flurtamone potentially can be used to control Italian ryegrass in winter wheat and barley, but not in oats. In general, wheat and oat injury with flurtamone was PPI > PRE ≥ early POST, particularly at 1.1 kg ha-1.

Field experiments evaluated barnyardgrass, broadleaf signalgrass, and rhizomatous johnsongrass control with sethoxydim, clethodim, fluazifop-P, quizalofop-P, and fenoxaprop applied alone and with bentazon or chlorimuron when crop oil concentrate or BCH 815 was used. Bentazon strongly antagonized sethoxydim and clethodim. BCH 815 partially reduced the antagonism with sethoxydim and clethodim compared with crop oil concentrate. Chlorimuron was most antagonistic towards fluazifop-P, fenoxaprop, and quizalofop-P, and BCH 815 did not reduce antagonism. Additional experiments evaluated Italian ryegrass control with diclofop applied alone and with bromoxynil, metribuzin, 2,4-D amine, or tribenuron plus thifensulfuron with these adjuvants. The amine formulation of 2,4-D and tribenuron plus thifensulfuron reduced Italian ryegrass control with diclofop, irrespective of the adjuvant.

The objective of this paper was to evaluate the effect of preplant applications of trifluralin on barley stand and yield, and control of grass weeds in field experiments during 1992 and 1993. Factors examined were: (1) crop planting patterns (conventional drill with rows 15 cm apart and deep-seeder drill with rows 25 cm apart), (2) herbicide application times (22 d before sowing and immediately before sowing), and (3) herbicide application. During 1993, hand-weeded plots also were established. Trifluralin applied preplant at 528 g ai/ha reduced weed density and biomass. Weed control was higher under conventional planting than under the deep planting pattern, and there was no effect of the time of application on herbicide efficacy. There was no herbicide injury to the crop, and grain yield was higher in treated than in untreated plots due to successful weed control.

Three approaches to data analysis were compared to describe competitive interactions between wheat and Italian ryegrass. Replacement series were performed using the two species at total densities of 100, 200, and 400 plants/ m2, and separate monoculture experiments for each species at densities from 33 to 800 plants/m2. Approaches to data analysis included: 1) conventional analysis of replacement series experiments, 2) development of synthetic no-interaction responses from monoculture experiments for comparison with results from mixtures, and 3) responses of the reciprocal yield of individual plants to variation in densities of the two species. Wheat was the superior competitor to ryegrass; however, the three approaches varied in ability to quantify this competitive relationship. The conventional replacement series analysis was least sensitive in describing the influences of either density or proportion on the plant association. The synthetic no-interaction approach provided the most detailed analysis of the influence of proportion on the species interaction. The reciprocal yield approach provided the simplest and most sensitive analysis of the joint influences of density and proportion. The latter approach also provided the most quantitative analysis of the influence of density on the species interaction. Plant density and species proportion are important variables for interpreting the process of plant competition.

Jointed goatgrass (Aegilops cylindrica) is a serious problem for winter wheat producers throughout the western U.S. Interference from this weed can severely reduce grain yield and contaminate harvested grain, resulting in dockage losses. There are currently no selective herbicides registered for controlling jointed goatgrass in wheat. Imazamox, an imidazolinone herbicide, was applied to an imidazolinone herbicide-resistant (IMI) wheat mutant of the winter wheat cultivar ‘Fidel.’ Jointed goatgrass control from spring postemergence application of imazamox ranged from 61 to 97% when applied at 36 g/ha. Negligible crop injury from imazamox treatment was observed at 36 g/ha at several locations under dryland environments in the U.S. Pacific Northwest. Wheat yield was increased 19 to 41% by imazamox treatment in three of four experiments. Percent dockage resulting from jointed goatgrass spikelet contamination of harvested wheat grain was eliminated by imazamox treatment. Introduction of the IMI trait into commercial wheat cultivars could provide an effective method for selective control of jointed goatgrass in winter wheat.

In most cases, a herbicide must traverse the cell wall, the plasma membrane, and organellar membranes of a plant cell to reach its site of action where accumulation causes phytotoxicity. The physicochemical characteristics of the herbicide molecule including lipophilicity and acidity, the plant cell membranes, and the electrochemical potential in the plant cell control herbicide absorption and accumulation. Most herbicides move across plant membranes via nonfacilitated diffusion because the membrane's lipid bilayer is permeable to neutral, lipophilic xenobiotics. Passive absorption of lipophilic, ionic herbicides or weak acids can be mediated by an ion-trapping mechanism where the less lipophilic, anionic form accumulates in alkaline compartments of the plant cell. A model that includes the pH and electrical gradients across plant cell membranes better predicts accumulation concentrations in plant cells of weak acid herbicides compared to a model that uses pH only. Herbicides also may accumulate in plant cells by conversion to nonphytotoxic metabolites, binding to cellular constituents, or partitioning into lipids. Evidence exists for herbicide transport across cell membranes via carrier-mediated processes where herbicide accumulation is energy dependent; absorption is saturable and slowed by metabolic inhibitors and compounds of similar structure.

Studies were conducted to determine whether cross-seeding of winter wheat, while maintaining an equal seeding rate per hectare, would increase wheat grain yields and help the wheat compete more effectively against Italian ryegrass than conventional seeding. Wheat was seeded at rates of 50, 100, and 150 kg ha-1 in conventional parallel rows or cross seeded in a grid pattern. Ryegrass was seeded at densities of 0, 20, 100, and 200 plants per m-2. Cross-seeded wheat yielded less grain than single-seeded wheat in 1990 in the absence of ryegrass; there were no differences in 1991. Cross seeding did not reduce competition from Italian ryegrass. Ryegrass was highly competitive against wheat, especially in 1990. Even 20 plants per m-2 reduced grain yield by an average of 38%. Wheat seeding rate had essentially no effect on grain yield in 1990, but the higher seeding rates reduced yields in 1991 because of extensive lodging. In 1991, wheat grain yields were higher in the cross-seeded plots than the single-seeded plots at the two high wheat seeding rates and highest ryegrass densities. Cross seeding does not appear promising as an aid to controlling ryegrass in Pacific Northwest wheat.

In field experiments, chlorsulfuron and CGA 131036 did not injure winter wheat. Both herbicides applied postemergence at 9 and 10 g ai/ha completely controlled flixweed. CGA 131036 controlled cutleaf eveningprimrose 81%, and chlorsulfuron controlled 91%. At similar rates, CGA 131036 was also less effective than chlorsulfuron on henbit and Italian ryegrass. Both controlled wild buckwheat 77% when applied postemergence at 18 g/ha. Neither of the sulfonylurea herbicides controlled prostrate spurge or woolly croton as effectively as 2,4-D. In a bean root bioassay, root growth inhibition by both herbicides was partially alleviated by adding isoleucine and valine.

Cotton and pitted morningglory emergence and dry weight decreased approximately 60 to 80% when these plants were grown under greenhouse conditions in the presence of increasing amounts (0.8 to 6.7 mg debris/g soil) of field-grown crimson clover or hairy vetch debris incorporated into the soil medium. Conversely, corn dry weight increased 20 to 75% when legume debris was placed on the soil surface; incorporated debris had very little effect on corn emergence or dry weight. Germination and seedling growth of corn, Italian ryegrass, cotton, pitted morningglory, and wild mustard decreased progressively, with species-dependent variation, when exposed to increasing concentrations (8.3 to 33.3 g debris/L) of aqueous crimson clover and hairy vetch extract. Mustard and ryegrass germination and growth were almost completely inhibited by full-strength extracts of both legumes. Bioassay species exhibited greater phytotoxic responses to hairy vetch than to crimson clover in the debris and extract studies. Emergence and growth of corn and cotton were not affected when planted into soil samples, containing root biomass and possible leaf and root exudates, collected from beneath field-grown hairy vetch and crimson clover plants. However, morningglory dry weight increased 35% in the presence of either legume root debris and accompanying soil.

A greenhouse experiment compared the vegetative growth in pure cultures and mixtures of winter Triticum aestivum cultivar ‘Mit’ and Lolium multiflorum cultivar ‘Marshall’ in continuously watered controls and drought treatments. Control L. multiflorum in pure culture 14 wk after planting produced more leaf area, tillers, and dry weights of stem and root than control T. aestivum in pure culture. The greater seed size, larger initial leaf area, and height allowed T. aestivum to produce greater final leaf area and dry stem weight in control mixtures than L. multiflorum. Watering following drought shifted the relative performance of the two species in pure cultures and mixtures compared to controls. The ability of T. aestivum to maintain a greater leaf expansion rate during drought and a greater leaf area afterward than L, multiflorum allowed T. aestivum to attain greater growth than L. multiflorum in pure cultures exposed to temporary drought followed by watering. Conversely, drought and its relief enhanced the relative competitiveness of L. multiflorum compared to controls in mixtures with T. aestivum. During 4 wk of watering following the drought, L. multiflorum in mixtures grew vigorously and was similar to T. aestivum in all measures except in height and dry stem weight. Thus, L. multiflorum was similar in root growth with T. aestivum in control and drought mixtures and had its aboveground competitiveness amplified by the cycle of drought and watering in this study. There was no evidence of an allelopathic interaction between the two species.

Ryegrass (Lolium multiflorum Lam. # LOLMU) control in wheat (Triticum aestivum L.) using chlorsulfuron {2-chloro-N-[[(4-methoxy-6-methyl-1,3,5-triazin-2-yl) amino] carbonyl] benzenesulfonamide} at 35 g ai/ha was significantly higher when applied preemergence (pre) than early postemergence (poe) to ryegrass with two to four leaves and wheat with three to five tillers. Metribuzin [4-amino-6-(1,1-dimethylethyl)-3-(methylthio)-1,2,4-triazin-5(4H)-one] applied at 420 g ai/ha early poe controlled more than 95% of the ryegrass, but control was unacceptable when the same rate was applied late poe when the ryegrass was at midtiller and wheat at full tiller. Wheat leaf burn, stunting, and stand reduction occurred where metribuzin was applied early poe but yields were not affected. Ryegrass control was excellent where diclofop {(±)-2-[4-(2,4-dichlorophenoxy)phenoxy] propanoic acid} was applied early poe at 560 g ai/ha. Wheat yields, averaged over years, for early poe applications of chlorsulfuron at 35 g/ha, metribuzin at 280 and 420 g/ha, and diclofop at 560 g/ha were 22, 22, 24, and 20% higher, respectively, than the untreated stands. Injury associated with the herbicide treatments was not observed on soybeans [Glycine max (L.) Merr.] planted following wheat.

Metolachlor, applied to the surface of leaching columns as dicalcium phosphate tablets, was recovered over a period of 25 d compared with 2 and 3 d for emulsifiable concentrate and granular formulations, respectively. In similar studies, approximately 27% of napropamide was recovered from tablets during the same period. Adding an adjuvant to the napropamide tablet mixture increased the recovery rate to 55%. Tablet hardness and size had little effect on the release rate of napropamide. Tablets containing 4% ai napropamide released greater amounts of herbicide at a single collection period than did tablets containing 1% ai. Adding an adjuvant to the napropamide tablet mixture significantly improved Italian ryegrass control in the area adjacent to herbicide placement.