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Echinoderms from the Ordovician of the Girvan district, Scotland, with the description of two new species

Published online by Cambridge University Press:  03 November 2011

Gerhard Regnéll
Affiliation:
Department of Historical Geology and Palaeontology, University of Lund, Sölvegatan 13, S-223 62 Lund, Sweden
C. R. C. Paul
Affiliation:
Department of Geology, University of Liverpool, P.O. Box 147, Liverpool L69 3BX, England

Abstract

Two new species of Ordovician echinoderms are described: Pleurocystites mitratus from the Kiln Mudstones (Caradoc) at Craighead, and Homocystites beggi from the Starfish Bed (Ashgill) of Threave Glen. Description of the former necessitated a review of the composition and status of the family Pleurocystitidae. The genus Dipleurocystis Jaekel 1918 and the subgenera Pleurocystites s.s. and Pleurocystella Paul, 1967 are rejected. The number and type of pectinirhombs; size of the periproct, composition of its border and nature of its covering plates (periproctals), and the ornament of the thecal plates and proximal columnals, are the most useful taxonomic criteria within the Pleurocystitidae. Pleurocystites mitratus is characterized by three conjunct pectinirhombs, a symmetrical theca without a rectal lobe and small periproctals each with a central pit. Homocystites beggi is distinguished from the closely related H. constrictus by strong granular ornament on the thecal plates.

Other echinoderm remains include plates of a cheirocrinid cystoid from the Kiln Mudstones.

Type
Research Article
Copyright
Copyright © Royal Society of Edinburgh 1981

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References

Barrande, J. 1887. Systeme Silurien du centre de la Boheme. le partie: Recherches paleontologiques. 1. Classe des echinodermes. Ordre des Cystides. Prague: W. Waagen.Google Scholar
Bassler, R. S. 1938. Pelmatozoa Palaeozoica (Generum et Genotyporum index et bibliographica). FOSSILIUM CATALOGUS 1 ANIMALIA 83.Google Scholar
Bassler, R. S. & Moodey, M. W. 1943. Bibliographic and faunal index of Paleozoic pelmatozoan echinoderms. SPEC PAP GEOL SOC AM 45.Google Scholar
Bather, F. A. 1899. A phylogenetic classification of the Pelmatozoa. REP BR ASSOC ADV SCI 68, 916–23.Google Scholar
Bather, F. A. 1900. The Echinoderms. In Lankester, E. R.A treatise on Zoology, vol. 3. London: A. & C. Black.Google Scholar
Bather, F. A. 1913. Caradocian Cystidea from Girvan. TRANS R SOC EDINBURGH 49, 359530.CrossRefGoogle Scholar
Bather, F. A. 1928. The fossil and its environment. Q J GEOL SOC LONDON 84, lxi–xcviii.Google Scholar
Billings, E. 1854. On some new genera and species of Cystidea from the Trenton Limestone. CAN J 2, 215–8, 250–3.Google Scholar
Bolton, T. E. 1970. Echinodermata from the Ordovician (Pleurocystites, Cremacrinus) and Silurian (Hemicystites, Protaxocrinus, Macnamarastylus) of Lake Timiskaming Region, Ontario and Quebec. BULL GEOL SURV CAN 187, 5966.Google Scholar
Breimer, A. 1963. On a specimen of Pleurocystites (Cystoidea) from an erratic boulder near Westerhaar (Netherlands). PROC K NED AKAD WET B66, 296302.Google Scholar
Broadhead, T. W. 1974. Re-evaluation of the morphology of Amecystis laevis (Raymond). J PALEONTOL 48, 670–3.Google Scholar
Broadhead, T. W. 1978. The Ordovician cystoid, Amecystis cordifiormis Sinclair. CAN J EARTH SCI 15, 173–4.Google Scholar
Broadhead, T. W. & Strimple, H. L. 1975. Respiration in a vagrant Ordovician cystoid, Amecystis. PALEOBIOLOGY 1, 312–9.Google Scholar
Dehm, R. 1932. Cystoideen aus dem rheinischen Unterdevon. NEUS JAHRB MINERAL GEOL PALAONTOL BEIL BAND ABT B 69, 6393.Google Scholar
Farmanfarmaian, A. 1966. The respiratory physiology of echinoderms. In Boolootian, R. A. (ed.) Physiology of Echinodermata, 245265. New York: Wiley.Google Scholar
Greig, D. C. 1971. The South of Scotland, 3rd edn. British Regional Geology Edinburgh: Inst. Geol. Sci.Google Scholar
Jaekel, O. 1899. Stammesgeschichte der Pelmatozoen. 1. Thecoidea und Cystoidea. Berlin: Springer.Google Scholar
Jaekel, O. 1918. Phylogenie und System der Pelmatozoen. PALAONTOL Z 1, 1128.CrossRefGoogle Scholar
Kesling, R. V. 1968. Cystoids. In Moore, R. C. (ed.) Treatise on invertebrate paleontology. S. Echinodermata vol. 1, S85–S267. Lawrence, Kansas: University of Kansas and Geological Society of America.Google Scholar
Kirk, E. 1911. The structure and relationships of certain eleutherozoic Pelmatozoa. PROC U S NATL MUS 41, 1137.CrossRefGoogle Scholar
Miller, S. A. 1889. North American geology and palaeontology. Cincinnati.Google Scholar
Miller, S. A. & Gurley, W. F. E. 1895. Description of new species of Palaeozoic Echinodermata. BULL ILLINOIS STATE MUS NAT HIST 6.Google Scholar
Neumayr, M. 1889. Die Stamme des Thierreiches. Wirbellose Thiere. Vienna: F. Tempsky.Google Scholar
Nicholson, H. A. & Etheridge, R. Jr. 1880. A monograph of the Silurian fossils of the Girvan district in Ayrshire with special reference to those contained in the “Gray collection”. Fasc III. Edinburgh: Blackwood.Google Scholar
Parsley, R. L. 1970. Revision of the North American Pleurocystitidae (Rhombifera-Cystoidea). BULL AM PALEONTOL 58, 135213.Google Scholar
Paul, C. R. C. 1967. The functional morphology and mode of life of the cystoid Pleurocystites E. Billings, 1854. SYMP ZOOL SOC LONDON 20, 105–21.Google Scholar
Paul, C. R. C. 1972. CheirocyStella antiqua gen. et sp. nov. from the Lower Ordovician of western Utah, and its bearing on the evolution of the Cheirocrinidae (Rhombifera: Glyptocystitida). GEOL STUD BRIGHAM YOUNG UNIV 19, 1563.Google Scholar
Paul, C. R. C. 1974. Regulaecystis devonica, a new Devonian pleurocystitid cystoid from Devon. GEOL MAG 111, 349–52.CrossRefGoogle Scholar
Regnéll, G. 1948. Echinoderms (Hydrophoridea, Ophiocistia) from the Ordovician (Upper Skiddavian, 3c) of the Oslo Region. NORSK GEOL TIDSSKR 27, 1458.Google Scholar
Regnéll, G. 1951. Revision of the Caradocian-Ashgillian cystoid fauna of Belgium with notes on isolated pelmatozoan stem fragments. MEM INST R SCI NAT BELG 120, 147.Google Scholar
Regnéll, G. 1960. The Lower Palaeozoic echinoderm faunas of the British Isles and Balto-Scandia. PALAEONTOLOGY 2, 161–79.Google Scholar
Sinclair, G. W. 1945. An Ordovician faunule from Quebec. CAN FIELD NAT 59, 71–4.Google Scholar
Sinclair, G. W. 1948. Three notes on Ordovician cystoids. J PALEONTOL 22, 301–14.Google Scholar
Slocom, A. W. & Foerste, A. F. 1924. New echinoderms from the Maquoketa beds of Fayette County, Iowa. REP IOWA GEOL SURV 29, 315–82.CrossRefGoogle Scholar
Sprinkle, J. 1974. New rhombiferan cystoids from the Middle Ordovician of Nevada. J PALEONTOL 48, 1174–201.Google Scholar
Tripp, R. P. 1980a. Trilobites from the Ordovician Balclatchie and lower Ardwell groups of the Girvan district, Scotland. TRANS R SOC EDINBURGH: EARTH SCI 71, 123–45.Google Scholar
Tripp, R. P. 1980b. Trilobites from the Ordovician Ardwell Group of the Craighead Inlier, Girvan district, Scotland. TRANS R SOC EDINBURGH: EARTH SCI 71, 147–57.Google Scholar
Ubaghs, G. 1953. Classe des Crinoides. In Piveteau, J. (ed.) Traite de Paleontologie vol. 3, 658773. Paris: Masson & Cie.Google Scholar
Ulrich, E. O. & Kirk, E. 1921. Amecystis, a new genus of Ordovician Cystidea. PROC BIOL SOC WASH 34, 147–8.Google Scholar
Zittel, K. A. von, 1879. Handbuch der Palaeontologie. 1 Palaeozoologie 1. Fasc. 3 (Echinodermata), 309564. Munich: R. Oldenbourg.Google Scholar