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Theropod teeth from the basalmost Cretaceous of Anoual (Morocco) and their palaeobiogeographical significance

Published online by Cambridge University Press:  16 February 2009

F. KNOLL  and
J. I. RUIZ-OMEÑACA
F. KNOLL*
Affiliation:
Departamento de Paleobiología, Museo Nacional de Ciencias Naturales-CSIC, C/José Gutiérrez Abascal 2, E-28006 Madrid, Spain
J. I. RUIZ-OMEÑACA
Affiliation:
Museo del Jurásico de Asturias, Rasa de San Telmo, E-33328 Colunga, Spain
*
†Author for correspondence: mcnfk854@mncn.csic.es
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Abstract

The theropod teeth from the Berriasian (Early Cretaceous) site of Anoual (N Morocco) are described. The assemblage is important in that it comes from one of the very few dinosaur sites of this age globally and the only one for the whole of Gondwana. The theropod teeth from Anoual are morphologically diverse. Most of the material possibly belongs to the clade Dromaeosauridae, which would be an early occurrence for this taxon. The palaeogeographic position of Anoual enables it to provide data on the dispersal events that affected terrestrial faunas during Mesozoic times. A Laurasian influence is evidenced by the presence of Velociraptorinae and, on the whole, the theropod fauna from Anoual provides support for the existence of a trans-Tethyan passage allowing terrestrial faunal interchanges during Late Jurassic and/or earliest Cretaceous times. Additionally, Anoual records the existence of diminutive theropods. However, it cannot yet be determined whether the small size of the specimens is genetic or ontogenetic.

Keywords

dinosaurCretaceouspalaeogeographyAfricaEurope
Type
Original Article
Information
Geological Magazine , Volume 146 , Issue 4 , July 2009 , pp. 602 - 616
Copyright
Copyright © Cambridge University Press 2009

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References

Alifanov, V. R. & Sennikov, A. G. 2001. [On the discovery of dinosaur remains in Moscow region]. Doklady Akademii nauk 376, 73–5 (in Russian).Google Scholar
Antunes, M. T. & Sigogneau, D. 1992. La faune de petits dinosaures du Crétacé terminal portugais. Comunicações dos Serviços Geológicos de Portugal 78, 49–62.Google Scholar
Antunes, M. T. & Sigogneau-Russell, D. 1996. Le Crétacé terminal portugais et son apport au problème de l'extinction des dinosaures. Bulletin du Muséum national d'Histoire naturelle, Section C 4, 595–606.Google Scholar
Averianov, A. O., Martin, T. & Bakirov, A. A. 2005. Pterosaur and dinosaur remains from the Middle Jurassic Balabansai Svita in the northern Fergana Depression, Kyrgyzstan (Central Asia). Palaeontology 48, 135–55.CrossRefGoogle Scholar
Bakker, R. T., Williams, M. & Currie, P. J. 1988. Nanotyrannus, a new genus of pygmy tyrannosaur, from the latest Cretaceous of Montana. Hunteria 1, 1–26.Google Scholar
Barco, J. L. & Ruiz-Omeñaca, J. I. 2001 a. Primeros dientes de terópodos (Dinosauria, Saurischia) en la Formación Villar del Arzobispo (Tithónico-Berriasiense): yacimientos Cuesta Lonsal y Las Cerradicas 2 (Galve, Teruel). In XVII Jornadas de la Sociedad Española de Paleontología: los Fósiles y la Paleogeografía, y Simposios de los Proyectos del Programa Internacional de Correlación Geológica (UNESCO/IUGS) N° 410 y 421 (eds Meléndez, G., Herrera, Z., Delvene, G. & Azanza, B.), pp. 239–46. Zaragoza: Seminario de Paleontología de Zaragoza.Google Scholar
Barco, J. L. & Ruiz-Omeñaca, J. I. 2001 b. Primeros restos postcraneales de terópodo (Dinosauria, Saurischia) en la Formación Villar del Arzobispo (Tithónico-Berriasiense): un centro vertebral caudal del yacimiento Carretera (Galve, Teruel). In XVII Jornadas de la Sociedad Española de Paleontología: los Fósiles y la Paleogeografía, y Simposios de los Proyectos del Programa Internacional de Correlación Geológica (UNESCO/IUGS) N° 410 y 421 (eds Meléndez, G., Herrera, Z., Delvene, G. & Azanza, B.), pp. 247–54. Zaragoza: Seminario de Paleontología de Zaragoza.Google Scholar
Barco, J. L., Ruiz-Omeñaca, J. I., Canudo, J. I. & Cuenca-Bescós, G. 2004. Dinosaurios. In Guía del Parque Paleontológico de Galve (ed. Barco, J. L.), pp. 64–88. Zaragoza: Paleoymás.Google Scholar
Barsbold, R. 1983. [Carnivorous dinosaurs from the Cretaceous of Mongolia]. Trudy–Sovmestnaya Sovetsko-Mongol'skaya Paleontologicheskaya Ekspeditsiya 19, 5–119 (in Russian, with English summary).Google Scholar
Barsbold, R. & Osmólska, H. 1999. The skull of Velociraptor (Theropoda) from the Late Cretaceous of Mongolia. Acta Palaeontologica Polonica 44, 189–219.Google Scholar
Baszio, S. 1997. Systematic palaeontology of isolated dinosaur teeth from the latest Cretaceous of south Alberta, Canada. Courier Forschunginstitut Senckenberg 196, 33–77.Google Scholar
Brinkman, D. L., Cifelli, R. L. & Czaplewski, N. J. 1998. First occurrence of Deinonychus antirrhopus (Dinosauria: Theropoda) from the Antlers Formation (Lower Cretaceous: Aptian–Albian) of Oklahoma. Oklahoma Geological Survey Bulletin 146, 1–27.Google Scholar
Burnham, D. A., Derstler, K. L., Currie, P. J., Bakker, R. T., Zhou, Z. & Ostrom, J. H. 2000. Remarkable new birdlike dinosaur (Theropoda: Maniraptora) from the Upper Cretaceous of Montana. The University of Kansas Paleontological Contributions 13, 1–14.Google Scholar
Canudo, J. I., Cuenca-Bescós, G. & Ruiz-Omeñaca, J. I. 1997. Dinosaurios dromeosaúridos [sic] (Saurischia: Theropoda) en el Barremiense Superior (Cretácico Inferior) de Castellote, Teruel. Geogaceta 22, 39–42.Google Scholar
Canudo, J. I., Ruiz-Omeñaca, J. I., Aurell, M., Barco, J. L. & Cuenca-Bescós, G. 2006. A megatheropod tooth from the late Tithonian–middle Berriasian (Jurassic–Cretaceous transition) of Galve (Aragón, NE Spain). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 239, 77–99.CrossRefGoogle Scholar
Carrano, M. T., Sampson, S. D. & Forster, C. A. 2002. The osteology of Masiakasaurus knopfleri, a small abelisauroid (Dinosauria: Theropoda) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology 22, 510–34.CrossRefGoogle Scholar
Codrea, V., Smith, T., Dica, P., Folie, A., Garcia, G., Godefroit, P. & Van Itterbeeck, J. 2002. Dinosaur egg nests, mammals and other vertebrates from a new Maastrichtian site of the Haţeg Basin (Romania). Comptes Rendus Palevol 1, 173–80.CrossRefGoogle Scholar
Company, J., Pereda Suberbiola, X., Ruiz-Omeñaca, J. I. & Buscalioni, A. D. 2005. A new species of Doratodon (Crocodyliformes: Ziphosuchia) from the Late Cretaceous of Spain. Journal of Vertebrate Paleontology 25, 343–53.CrossRefGoogle Scholar
Currie, P. J. 1995. New information on the anatomy and relationships of Dromaeosaurus albertensis (Dinosauria: Theropoda). Journal of Vertebrate Paleontology 15, 576–91.CrossRefGoogle Scholar
Currie, P. J. & Azuma, Y. 2006. New specimens, including a growth series, of Fukuiraptor (Dinosauria, Theropoda) from the Lower Cretaceous Kitadani Quarry of Japan. Journal of the Paleontological Society of Korea 22, 173–93.Google Scholar
Currie, P. J. & Chen, P. J. 2001. Anatomy of Sinosauropteryx prima from Liaoning, northeastern China. Canadian Journal of Earth Sciences 38, 1705–27.CrossRefGoogle Scholar
Currie, P. J., Rigby, K. Jr & Sloan, R. E. 1990. Theropod teeth from the Judith River Formation of southern Alberta, Canada. In Dinosaur Systematics: Approaches and Perspectives (eds Carpenter, K. & Currie, P. J.), pp. 107–25. New York: Cambridge University Press.CrossRefGoogle Scholar
Currie, P. J. & Varricchio, D. J. 2004. A new dromaeosaurid from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada. In Feathered Dragons (eds Currie, P. J., Koppelhus, E. B., Shugar, M. A. & Wright, J. L.), pp. 112–32. Bloomington: Indiana University Press.Google Scholar
Dal Sasso, C. & Signore, M. 1998. Exceptional soft-tissue preservation in a theropod dinosaur from Italy. Nature 392, 383–7.CrossRefGoogle Scholar
Dercourt, J., Guetani, M., Vrielynk, B., Barrier, E., Biju-Duval, B., Brunet, M.-F., Cadet, J.-P., Crasquin, S. & Sandulescu, M. (eds) 2000. Peri-Tethys palaeogeographical Atlas-Explanatory Notes. Paris: Gauthier-Villars, 268 pp., 24 maps.Google Scholar
Dercourt, J., Ricou, L.-E. & Vrielynck, B. (eds) 1993. Atlas Tethys-Palaeoenvironmental Maps-Explanatory Notes. Paris: Gauthier-Villars, 307 pp., 14 maps.Google Scholar
Evans, S. E. & Milner, A. R. 1994. Middle Jurassic microvertebrate assemblages from the British Isles. In In the Shadow of the Dinosaurs: Early Mesozoic Tetrapods (eds Fraser, N. and Sues, H.-D..), pp. 303–21. Cambridge: Cambridge University Press.Google Scholar
Evans, S. E. & Sigogneau-Russell, D. 2001. A stem-group caecilian (Lissamphibia: Gymnophiona) from the Lower Cretaceous of North Africa. Palaeontology 44, 259–73.CrossRefGoogle Scholar
Fanti, F. & Therrien, F. 2007. Theropod tooth assemblages from the Late Cretaceous Maevarano Formation and the possible presence of dromaeosaurids in Madagascar. Acta Palaeontologica Polonica 52, 155–66.Google Scholar
Farlow, J. O., Brinkman, D. L., Abler, W. L. & Currie, P. J. 1991. Size, shape, and serration density of theropod dinosaur lateral teeth. Modern Geology 16, 161–98.Google Scholar
Galton, P. M. 1996. Notes on Dinosauria from the Upper Cretaceous of Portugal. Neues Jahrbuch für Geologie und Paläontologie Monatshefte 1996, 83–90.CrossRefGoogle Scholar
Garcia, G., Pincemaille, M., Vianey-Liaud, M., Marandat, B., Lorenz, E., Cheylan, G., Cappetta, H., Michaux, J. & Sudre, J. 1999. Découverte du premier squelette presque complet de Rhabdodon priscus (Dinosauria, Ornithopoda) du Maastrichtien inférieur de Provence. Comptes Rendus de l'Académie des Sciences de Paris, Sciences de la Terre et des Planètes 328, 415–21.Google Scholar
Gardner, J. D., Evans, S. E. & Sigogneau-Russell, D. 2003. New albanerpetontid amphibians from the Early Cretaceous of Morocco and Middle Jurassic of England. Acta Palaeontologica Polonica 48, 301–19.Google Scholar
Gauthier, J. A. 1986. Saurischian monophyly and the origin of birds. Memoirs of the California Academy of Sciences 8, 1–55.Google Scholar
Gheerbrant, E. & Rage, J.-C. 2006. Paleobiogeography of Africa: How distinct from Gondwana and Laurasia? Palaeogeography, Palaeoclimatology, Palaeoecology 241, 224–46.CrossRefGoogle Scholar
Göhlich, U. B. & Chiappe, L. M. 2006. A new carnivorous dinosaur from the Late Jurassic Solnhofen archipelago. Nature 440, 329–32.CrossRefGoogle ScholarPubMed
Goodwin, M. B., Clemens, W. A., Hutchison, J. H., Wood, C. B., Zavada, M. S., Kemp, A., Duffin, C. J. & Schaff, C. R. 1999. Mesozoic continental vertebrates with associated palynostratigraphic dates from the northwestern Ethiopian plateau. Journal of Vertebrate Paleontology 19, 728–41.CrossRefGoogle Scholar
Grigorescu, D. 1984. New paleontological data on the dinosaur beds from the Hateg Basin. In 75 years of the Laboratory of Paleontology (eds Petrescu, J. & Dragastan, O..), pp. 111–18. Bucureşti: Universitatea din-Bucureşti.Google Scholar
Haddoumi, H., Alméras, Y., Bodergat, A. M., Charrière, A., Mangold, C. & Benshili, K. 1998. Ages et environnements des couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc). Comptes Rendus de l'Académie des Sciences de Paris, Sciences de la Terre et des Planètes 327, 127–33.Google Scholar
Hahn, G. & Hahn, R. 2003. New multituberculate teeth from the Early Cretaceous of Morocco. Acta Palaeontologica Polonica 48, 349–56.Google Scholar
Hartman, S., Lovelace, D. & Wahl, W. 2005. Phylogenetic assessment of a maniraptoran from the Morrison Formation. Journal of Vertebrate Paleontology 25, 67A–68A.Google Scholar
Holtz, T. R. Jr. 1995. A new phylogeny of the Theropoda. Journal of Vertebrate Paleontology 15, 35A.Google Scholar
Holtz, T. R. Jr, Molnar, R. E. & Currie, P. J. 2004. Basal Tetanurae. In The Dinosauria, Second Edition (eds Weishampel, D. B., Dodson, P. & Osmólska, H..), pp. 71–110. Berkeley: University of California Press.Google Scholar
Holtz, T. R. Jr & Osmólska, H. 2004. Saurischia. In The Dinosauria, Second Edition (eds Weishampel, D. B., Dodson, P. & Osmólska, H..), pp. 21–4. Berkeley: University of California Press.Google Scholar
von Huene, F. 1914. Das natürliche System der Saurischia. Zentralblatt für Mineralogie, Geologie und Paläontologie, Abteilung B 1914, 154–8.Google Scholar
Hwang, S. H., Norell, M. A., Ji, Q. & Gao, K. 2002. New specimens of Microraptor zhaoianius (Theropoda: Dromaeosauridae) from northeastern China. American Museum Novitates 3381, 1–44.2.0.CO;2>CrossRefGoogle Scholar
Hwang, S. H., Norell, M. A., Ji, Q. & Gao, K. 2004. A large compsognathid from the Early Cretaceous Yixian Formation of China. Journal of Systematic Palaeontology 2, 13–30.CrossRefGoogle Scholar
Jones, M. E. H., Evans, S. E. & Sigogneau-Russell, D. 2003. Early Cretaceous frogs from Morocco. Annals of Carnegie Museum 72, 65–97.CrossRefGoogle Scholar
Knoll, F. 2000. Pterosaurs from the Lower Cretaceous (?Berriasian) of Anoual, Morocco. Annales de Paléontologie 86, 157–64.CrossRefGoogle Scholar
Laurent, Y., Bilotte, M. & Le Loeuff, J. 2002. Late Maastrichtian continental vertebrates from southwestern France: correlation with marine fauna. Palaeogeography, Palaeoclimatology, Palaeoecology 187, 121–35.CrossRefGoogle Scholar
Leonardi, G. & Carvalho, I. de S. 2002. Icnofósseis da Bacia do Rio do Peixe, PB: o mais marcante registro de pegadas de dinossauros do Brasil. In Sítios geológicos e paleontológicos do Brasil (eds Schobbenhaus, C., Campos, D. de A., Queiroz, E. T., Winge, M. & Berbert-Born, M..), pp. 101–11. Brasilia: SIGEP.Google Scholar
Leonardi, G. & Carvalho, I. de S. 2007. Dinosaur ichnocoenosis from Sousa and Uiraúna-Brejo das Freiras Basins, northeast Brazil. In Paleontologia: Cenários de Vida, vol. 1 (eds Carvalho, I. de S., Cassab, R. de C. T., Schwanke, C., Carvalho, M. de A., Fernandes, A. C. S., Rodrigues, M. A. da C., Carvalho, M. S. S. de, Arai, M. & Oliveira, M. E. Q..), pp. 355–69. Rio de Janeiro: Interciência.Google Scholar
Lindgren, J., Currie, P. J., Rees, J., Siverson, M., Lindström, S., Alwmark, C. 2008. Theropod dinosaur teeth from the lowermost Cretaceous Rabekke Formation on Bornholm, Denmark. Geobios 41, 253–62.CrossRefGoogle Scholar
Maganuco, S., Cau, A. & Pasini, G. 2005. First description of theropod remains from the Middle Jurassic (Bathonian) of Madagascar. Atti della Società Italiana di Scienze Naturali e del Museo civico di Storia Naturale di Milano 146, 165–202.Google Scholar
Makovicky, P. J., Apesteguía, S. & Agnolín, F. L. 2005. The earliest dromaeosaurid theropod from South America. Nature 437, 1007–11.CrossRefGoogle ScholarPubMed
Makovicky, P. & Norell, M. A. 2004. Troodontidae. In The Dinosauria, Second Edition (eds Weishampel, D. B., Dodson, P. and Osmólska, H..), pp. 184–95. Berkeley: University of California Press.Google Scholar
Marsh, O. C. 1881. Principal characters of American Jurassic dinosaurs; Part V. American Journal of Science 21, 417–23.CrossRefGoogle Scholar
Mateer, N. J. 1987. A new report of a theropod dinosaur from South Africa. Palaeontology 30, 141–5.Google Scholar
Matthew, W. D. & Brown, B. 1922. The family Deinodontidae, with notice of a new genus from the Cretaceous of Alberta. Bulletin of the American Museum of Natural History 46, 367–85.Google Scholar
Milner, A. 2002. Theropod dinosaurs of the Purbeck Limestone Group, Southern England. Special Papers in Palaeontology 68, 191–201.Google Scholar
Mojon, P. O., Haddoumi, H. & Charrière, A. 2005. Nouvelles données sur les Charophytes et Ostracodes du Jurassique–Crétacé de l'Atlas marocain. In Recherches sur la Pangée mésozoïque (ed. Mojon, P. O..), pp. 4–47. Lausanne: Digit Presse.Google Scholar
Norell, M. A., Clark, J. M., Dashzeveg, D., Barsbold, R., Chiappe, L. M., Davidson, A. R., McKenna, M. C., Perle, A. & Novacek, M. J. 1994. A theropod dinosaur embryo and the affinities of the Flaming Cliffs dinosaur eggs. Science 266, 779–82.CrossRefGoogle ScholarPubMed
Norell, M. A., Clark, J. M., Turner, A. H., Makovicky, P. J., Barsbold, R. & Rowe, T. 2006. A new dromaeosaurid theropod from Ukhaa Tolgod (Ömnögov, Mongolia). American Museum Novitates 3545, 1–51.CrossRefGoogle Scholar
Novas, F. E. & Pol, D. 2005. New evidence on deinonychosaurian dinosaurs from the Late Cretaceous of Patagonia. Nature 433, 858–61.CrossRefGoogle ScholarPubMed
Osborn, H. F. 1924. Three new Theropoda, Protoceratops zone, central Mongolia. American Museum Novitates 144, 1–12.Google Scholar
Ostrom, J. H. 1969. Osteology of Deinonychus antirrhopus, an unusual theropod from the Lower Cretaceous of Montana. Peabody Museum of Natural History Bulletin 30, 1–165.Google Scholar
Ostrom, J. H. 1978. The osteology of Compsognathus longipes Wagner. Zitteliana 4, 73–118.Google Scholar
Ostrom, J. H. 1990. Dromaeosauridae. In The Dinosauria (eds Weishampel, D. B., Dodson, P. & Osmólska, H..), pp. 269–79. Berkeley: University of California Press.Google Scholar
Owen, R. 1842. Report on British fossil reptiles; Part II. Report of the British Association for the Advancement of Science 1841, 60–204.Google Scholar
Paul, G. S. 1988 a. The small predatory dinosaurs of the mid-Mesozoic: the horned theropods of the Morrison and Great Oolite – Ornitholestes and Proceratosaurus – and the sickle-claw theropods of the Cloverly, Djadokhta and Judith River – Deinonychus, Velociraptor and Saurornitholestes. Hunteria 2, 1–9.Google Scholar
Paul, G. S. 1988 b. Predatory Dinosaurs of the World. New York: Simon and Schuster, 464 pp.Google Scholar
Pereda Suberbiola, X. 1999. Las faunas finicretácicas de dinosaurios ibéricos. Zubía 17, 259–79.Google Scholar
Pol, C., Buscalioni, A. D., Carballeira, J., Francés, V., López Martinez, N., Marandat, B., Moratalla, J. J., Sanz, J. L., Sigé, B. & Villatte, J. 1992. Reptiles and mammals from the Late Cretaceous new locality Quintanilla del Coco (Burgos Province, Spain). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 184, 279–314.Google Scholar
Prieto-Márquez, A., Gaete, R., Galobart, A. & Ardèvol, L. 2000. A Richardoestesia-like theropod tooth from the Late Cretaceous foredeep, south-central Pyrenees, Spain. Eclogae Geologicae Helvetiae 93, 497–501.Google Scholar
Rauhut, O. W. M. 2000. The dinosaur fauna from the Guimarota mine. In Guimarota – A Jurassic Ecosystem (eds Martin, T. & Krebs, B..), pp. 75–82. München: Dr Friedrich Pfeil.Google Scholar
Rauhut, O. W. M. 2001. Herbivorous dinosaurs from the Late Jurassic (Kimmeridgian) of Guimarota, Portugal. Proceedings of the Geologists' Association 112, 275–83.CrossRefGoogle Scholar
Rauhut, O. W. M. 2002. Dinosaur teeth from the Barremian of Uña, Province of Cuenca, Spain. Cretaceous Research 23, 255–63.CrossRefGoogle Scholar
Rauhut, O. W. M. 2004. Provenance and anatomy of Genyodectes serus, a large-toothed ceratosaur (Dinosauria: Theropoda) from Patagonia. Journal of Vertebrate Paleontology 24, 894–902.CrossRefGoogle Scholar
Rauhut, O. W. M. 2005. Osteology and relationships of a new theropod dinosaur from the Middle Jurassic of Patagonia. Palaeontology 48, 87–110.CrossRefGoogle Scholar
Rauhut, O. W. M. & Werner, C. 1995. First record of the family Dromaeosauridae (Dinosauria: Theropoda) in the Cretaceous of Gondwana (Wadi Milk Formation, northern Sudan). Paläontologische Zeitschrift 69, 475–89.CrossRefGoogle Scholar
Rauhut, O. W. M. & Zinke, J. 1995. A description of the Barremian dinosaur fauna from Uña with a comparison to that of Las Hoyas. In II International Symposium on Lithographic Limestones, Lleida-Cuenca (Spain), 9th–16th July 1995 – Extended Abstracts (ed. Meléndez, N..), pp. 123–6. Cantoblanco: Universidad Autónoma de Madrid.Google Scholar
Royo-Torres, R., Cobos, A., Alcalá, L. & Bello, Y. 2003. Primeros restos de dinosaurio en el Cretácico Inferior de Riodeva (Teruel). In XIX Jornadas de la Sociedad Española de Paleontología, Morella, 16/18 Octubre 2003 – Libro de Resúmenes (eds Alonso, M. V. Pardo & Gozalo, R..), p. 147. Morella: Ajuntament de Morella.Google Scholar
Ruiz-Omeñaca, J. I. & Canudo, J. I. 2001. Dos yacimientos excepcionales con vertebrados continentales del Barremiense (Cretácico Inferior) de Teruel: Vallipón y La Cantalera. Naturaleza aragonesa 8, 8–17.Google Scholar
Ruiz-Omeñaca, J. I. & Canudo, J. I. 2003. Un nuevo dinosaurio terópodo (“Prodeinodon” sp.) en el Cretácico Inferior de la Cantalera (Teruel). Geogaceta 34, 111–14.Google Scholar
Ruiz-Omeñaca, J. I., Canudo, J. I. & Cuenca-Bescós, G. 1996. Dientes de dinosaurios (Ornithischia y Saurischia) del Barremiense superior (Cretácico inferior) de Vallipón (Castellote, Teruel). Mas de las Matas 15, 59–103.Google Scholar
Ruiz-Omeñaca, J. I., Canudo, J. I., Cuenca-Bescós, G. & Amo, O. 1998. Theropod teeth from the Lower Cretaceous of Galve (Teruel, Spain). In Third European Workshop of Vertebrate Palaeontology: Programme and Abstracts, Field Guide (eds Jagt, J. W. M., Lambers, P. H., Mulder, E. W. A. & Schulp, A. S..), pp. 62–3. Maastricht: Natuurhistorisch Museum Maastricht.Google Scholar
Samman, T., Powell, G. L., Currie, P. J. & Hills, L. V. 2005. Morphometry of the teeth of western North American tyrannosaurids and its applicability to quantitative classification. Acta Palaeontologica Polonica 50, 757–76.Google Scholar
Sankey, J. T. 2008. Diversity of Latest Cretaceous (Late Maastrichtian) small theropods and birds: teeth from the Lance and Hell Creek Formations, USA. In Vertebrate Microfossil Assemblages: their role in paleoecology and paleobiogeography (eds by Sankey, J. T. & Baszio, S..), pp. 117–34. Bloomington: Indiana University Press.Google Scholar
Sankey, J. T., Brinkman, D. B., Guenther, M. & Currie, P. J. 2002. Small theropod and bird teeth from the Late Cretaceous (Late Campanian) Judith River Group, Alberta. Journal of Paleontology 76, 751–63.2.0.CO;2>CrossRefGoogle Scholar
Sankey, J. T., Standhardt, B. R. & Schiebout, J. A. 2005, Theropod teeth from the Upper Cretaceous (Campanian–Maastrichtian), Big Bend National Park, Texas. In The Carnivorous Dinosaurs (ed. Carpenter, K..), pp. 127–52. Bloomington: Indiana University Press.Google Scholar
Seeley, H. G. 1888. On the classification of the fossil animals commonly named Dinosauria. Proceedings of the Royal Society of London 43, 165–71.Google Scholar
Sereno, P. C. & Brusatte, S. L. 2008. Basal abelisaurid and carcharodontosaurid theropods from the Lower Cretaceous Elrhaz Formation of Niger. Acta Palaeontologica Polonica 53, 15–46.CrossRefGoogle Scholar
Sereno, P. C., Dutheil, D. B., Iarochene, M., Larsson, H. C. E., Lyon, G. H., Magwene, P. M., Sidor, C. A., Varricchio, D. J. & Wilson, J. A. 1996. Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation. Science 272, 986–91.CrossRefGoogle ScholarPubMed
Sigogneau-Russell, D. 1999 a. Réévaluation des Peramura (Mammalia, Cladotheria) sur la base de nouveaux spécimens du Crétacé inférieur d'Angleterre et du Maroc. Geodiversitas 21, 93–127.Google Scholar
Sigogneau-Russell, D. 1999 b. Le site d'Anoual et sa riche faune de microvertébrés. In Maroc, Mémoire de la Terre (ed. Blandin, P..), pp. 148–51. Paris: Muséum national d'Histoire naturelle.Google Scholar
Sigogneau-Russell, D. 2003. Diversity of triconodont mammals from the Early Cretaceous of North Africa – affinities of the amphilestids. Palaeovertebrata 32, 27–55.Google Scholar
Sigogneau-Russell, D., Evans, S. E., Levine, J. F. & Russell, D. A. 1998. The Early Cretaceous microvertebrate locality of Anoual, Morocco: a glimpse at the small vertebrate assemblages of Africa. New Mexico Museum of Natural History and Science Bulletin 14, 177–81.Google Scholar
Sigogneau-Russell, D., Monbaron, M. & Kaenel, de E. 1990. Nouvelles données sur le gisement à Mammifères mésozoïques du Haut–Atlas marocain. Geobios 23, 461–83.CrossRefGoogle Scholar
Smith, J. B. 2005. Heterodonty in Tyrannosaurus rex: implications for the taxonomic and systematic utility of theropod dentitions. Journal of Vertebrate Paleontology 25, 865–87.CrossRefGoogle Scholar
Smith, J. B. 2007. Dental morphology and variation in Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar. Society of Vertebrate Paleontology Memoir 8, 103–26.CrossRefGoogle Scholar
Smith, J. B. & Dodson, P. 2003. A proposal for a standard terminology of anatomical notation and orientation in fossil vertebrate dentitions. Journal of Vertebrate Paleontology 23, 1–14.CrossRefGoogle Scholar
Smith, J. B., Vann, D. R. & Dodson, P. 2005. Dental morphology and variation in theropod dinosaurs: implications for the taxonomic identification of isolated teeth. The Anatomical Record Part A: Discoveries in Molecular, Cellular, and Evolutionary Biology 285A, 699–736.CrossRefGoogle Scholar
Stromer, E. 1934. Die Zähne des Compsognathus und Bemerkungen über dass Gebiss der Theropoda. Centralblatt für Mineralogie, Geologie und Paläontologie Abteilung B 1934, 74–85.Google Scholar
Sues, H.-D. 1978. A new small theropod dinosaur from the Judith River Formation (Campanian) of Alberta, Canada. Zoological Journal of the Linnean Society 62, 381–400.CrossRefGoogle Scholar
Suñer, M., Santisteban, C. de & Galobart, A. 2005. Nuevos restos de Theropoda del Jurásico Superior–Cretácico Inferior de la Comarca de los Serranos (Valencia). Revista Española de Paleontología, Número extraordinario 10, 93–9.Google Scholar
Sweetman, S. C. 2004. The first record of velociraptorine dinosaurs (Saurischia, Theropoda) from the Wealden (Early Cretaceous, Barremian) of southern England. Cretaceous Research 25, 353–64.CrossRefGoogle Scholar
Torices Hernández, A. 2002. Los dinosaurios terópodos del Cretácico superior de la Cuenca de Tremp (Pirineos Sur-Centrales, Lleida). Coloquios de Paleontología 53, 139–46.Google Scholar
Torices Hernández, A. 2003. Estudio preliminar de dientes aislados de terópodos en el Cretácico superior de la Cuenca de Tremp (Pirineos Sur-Centrales, Lleida). In II Jornadas internacionales sobre Paleontología de Dinosaurios y su Entorno (eds Vidarte, C. Fuentes & Fernández-Baldor, F. Torcida.), pp. 213–20. Salas de los Infantes: Colectivo Arqueológico y Paleontológico de Salas.Google Scholar
Torices, A., Ruiz-Omeñaca, J. I., Canudo, J. I. & López-Martínez, N. 2004. Nuevos datos sobre los dinosaurios terópodos (Saurischia: Theropoda) del Cretácico superior de los Pirineos Sur-Centrales (Huesca y Lleida). Geo-Temas 6, 71–4.Google Scholar
Turner, A. H., Hwang, S. H. & Norell, M. A. 2007. A small derived theropod from Öösh, Early Cretaceous, Baykhangor Mongolia. American Museum Novitates 3557, 1–27.CrossRefGoogle Scholar
Turner, A. H., Pol, D., Clarke, J. A., Erickson, G. M. &. Norell, M. A. 2007. A basal dromaeosaurid and size evolution preceding avian flight. Science 317, 1378–81.CrossRefGoogle ScholarPubMed
Valença, L. M. M., Neumann, V. H. & Mabesoone, J. M. 2003. An overview on Callovian–Cenomanian intracratonic basins of Northeast Brazil: onshore stratrigraphic record of the opening of the southern Atlantic. Geologica Acta 1, 261–75.Google Scholar
Wang, S. C. & Dodson, P. 2006. Estimating the diversity of dinosaurs. Proceedings of the National Academy of Sciences of the United States of America 103, 13601–5.CrossRefGoogle ScholarPubMed
Weishampel, D. B., Barrett, P. M., Coria, R. A., Le Loeuff, J., Xu, X., Zhao, X.-J., Sahni, A., Gomani, E. M. P. & Noto, C. R. 2004. Dinosaur distribution. In The Dinosauria, Second Edition (eds Weishampel, D. B., Dodson, P. & Osmólska, H..), pp. 517–606. Berkeley: University of California Press.Google Scholar
Werner, C. 1995. Neue Funde von mesozoischen Wirbeltieren in Äthiopien. Berliner Geowissenschaftliche Abhandlungen 16, 377–83.Google Scholar
Xu, X. & Wang, X. 2004. A new dromaeosaur (Dinosauria: Theropoda) from the Early Cretaceous Yixian Formation of western Liaoning. Vertebrata PalAsiatica 42, 111–19.Google Scholar
Xu, X. & Wu, X. 2001. Cranial morphology of Sinornithosaurus millenii Xu et al. 1999 (Dinosauria: Theropoda: Dromaeosauridae) from the Yixian Formation of Liaoning, China. Canadian Journal of Earth Sciences 38, 1739–52.CrossRefGoogle Scholar
Xu, X., Zhou, Z.-H. & Wang, X.-L. 2000. The smallest known non-avian theropod dinosaur. Nature 408, 705–8.CrossRefGoogle ScholarPubMed
Zinke, J. 1998. Small theropod teeth from the Upper Jurassic coal mine of Guimarota (Portugal). Paläontologische Zeitschrift 72, 179–89.CrossRefGoogle Scholar