The Andean orogeny in the Patagonian Cordillera of southern South America reflects the consequences of the Mesozoic and Cenozoic subduction of an oceanic plate beneath the South American continental margin. The geological evolution of the region has been influenced by the Eocene collision and subduction of the Farallon–Aluk Ridge and the Miocene–Recent subduction of the Chile Ridge. Another aspect of plate interaction during this period was two intervals of rapid plate convergence, one at 50–42 Ma, and the other at 25–10 Ma, between the South American and the oceanic plates. It has been proposed that the collision of the Chile Ridge with the trench was responsible for the development, at least in part, of the Patagonian fold and thrust belt. This belt extends for more than 1000 km along the eastern foothills of the southern Andes between 46° and 54° S along the southwestern rim of the Austral Basin. The interpretation of a link between subduction of the ridge and formation of the fold and thrust belt is based on assumed time coincidences between contractional tectonism and the collision of ridge segments during Middle and Late Miocene times. The main Tertiary contractional events in the Patagonian fold and thrust belt took place during latest Cretaceous–Palaeocene–Eocene and during Miocene times. Although the timing of deformation is still poorly constrained, the evidence currently available suggests that there is little or no relationship between the timing of the fold and thrust belt and the collision of ridge segments. Most if not all of the contractional tectonism pre-dated the latest episodes of ridge collision. Collision of a ridge crest with the continental margin has been active for the past 14 to 15 million years. Contrary to the suggestion of a relationship between ridge subduction and compression, the main result of this collision has been fast uplift and extensional tectonism. The initiation of the Patagonian fold and thrust belt in latest Cretaceous or early Tertiary times coincided with a fundamental change in the tectonic evolution of the Austral Basin. Throughout the Cretaceous most of this basin subsided as a broad backarc continental shelf. Only in latest Cretaceous times, and coinciding with the initiation of the fold and thrust belt, the basin underwent a transition to a retro-arc foreland basin. This change to an asymmetrically subsiding foreland basin, with an associated foreland fold and thrust belt, was related to uplift of the Andean orogenic belt in the west.

The Bossòst dome is an E–W-trending elongated structural and metamorphic dome developed in Cambro-Ordovician metasedimentary rocks in the Variscan Axial Zone of the central Pyrenees. A steep fault separates a northern half-dome, cored by massif granite, from an E–W-trending doubly plunging antiform with granitic sills and dykes in the core to the south. The main foliation is a flat-lying S1/2 schistosity that grades into a steeper-dipping slaty cleavage at the dome margins. Three major deformational and two metamorphic phases can be differentiated. S1/2 schistosity is an axial planar cleavage to W-vergent recumbent folding that probably occurred in mid-Westphalian time. Peak regional metamorphism M1 is characterized by static growth of staurolite and garnet following thermal relaxation of the previously thickened crust. Strong non-coaxial deformation recording uniform top-to-the-SE extension during D2a is preserved in staurolite–garnet schists in a 1.5 km thick, shallowly SE-dipping zone in the southeastern dome. A 500 m thick contact aureole (M2) was imprinted on the regionally metamorphosed rocks following the intrusion the Bossòst granite during D2b. More coaxial deformation prevailed during synkinematic growth of M2 phases in the inner part of the contact aureole around the northern part of the dome, where it obliterated D2a fabrics. Progressive non-coaxial deformation continued in the southeastern antiform and is recorded by late-synkinematic growth of cordierite. Successive overprinting of the M1 staurolite–garnet assemblage by andalusite and cordierite of M2 is preserved in the southern part. The assemblage muscovite+cordierite+staurolite+biotite is considered metastable, given the low Mn and Zn contents of staurolite and cordierite, and interpreted as the result of prograde metamorphism during decompression. P–T conditions during M2 were approximately 3 kbar and 600 °C. Pervasive crenulations and mesoscopic to regional southerly verging folds are the result of D3 NNE–SSW compression post-dating ductile deformation and contact metamorphism. Polymetamorphic assemblages of the Bossòst dome preserve a regionally confined zone of ESE-directed extensional shearing within an overall N–S compressional setting. Exact timing of extensional shearing is not known, but can be constrained by recumbent folding during the mid-Westphalian and granitic intrusions, which confine it to Late Carboniferous time (c. 305 Ma). Crustal-scale flat-lying extensional shear zones with similar orientation and time frame are observed in the Hospitalêt massif of the eastern Axial Zone. This suggests that crustal extension, though probably restricted by regional strain partitioning over orthogneiss or intruding granitic bodies within an overall compressive setting, was not uncommon in Late Carboniferous time in the Axial Zone of the Pyrenees.

The Troodos Massif of Cyprus exposes a classic and much-studied ophiolite sequence representing oceanic crust of Late Cretaceous age. K-Ar dating of the sheeted dykes and of the overlying pillow lavas gives a range of 83±3 Ma (earliest Campanian) and 75±5 Ma (late Campanian) respectively for the formation of the upper levels of the ophiolite. An autochthonous sequence of Late Cretaceous to Recent age sediments is exposed resting on the ophiolite, the oldest part of which reflects sedimentation in an apparently deep marine, oceanic setting. Little biostratigraphical information is available to constrain the chronostratigraphy of these sediments in relation to the complex geological history of the island, including the uplift and unroofing of the ophiolite, despite their richness in microfauna and flora. This paper provides an integrated biostratigraphical study based on radiolaria, planktonic foraminifera and calcareous nannofossils of the oldest part of the supra-ophiolite succession, of the Cretaceous Perapedhi, Kannaviou, Moni and Kathikas formations. For the first time, well-defined micropalaeontological evidence establishes the relative ages of these formations. The umbers of the Perapedhi formation are no younger than Campanian in age while the volcaniclastic sediments of the Kannaviou formation were also deposited during Campanian times. The matrix of the Moni formation contains a microfauna consistent with the hypothesis that it is derived from the Kannaviou. In contrast, the Kathikas formation is composed of sediment derived mainly from the allochthonous Mamonia complex, but autochthonous pelagic interbed horizons demonstrate that it was deposited probably within a very short time interval during the late Maastrichtian. Key index species are figured, including the first published micrographs of Cretaceous planktonic foraminifera from Cyprus.

1. Succession of Silurian Rocks in the South of Scotland.—The oldest fossiliferous strata yet detected in the Silurian district of the South of Scotland are the dark Graptolitic shales forming the well-known Moffat Series. The total thickness of this group is under 600 feet, but it is nevertheless capable of separation into three very distinct palæontological divisions, each characterized by a large proportion of peculiar fossils. The first of these divisions is of Llandeilo age, the second represents a portion of the Bala formation of North Wales, and the third, or Birkhill division, appertains, at least in part, to the epoch of the Lower Llandovery.

Plant fossils are described from the Cuche Formation, Eastern Cordillera, Colombia in the area of Floresta. Those identified as Colpodexylon cf. deatsii Banks and cf. Archaeopteris sp. suggest an earliest Late Devonian (Frasnian) age for the formation. These or similar taxa are also found in contemporaneous deposits in western Venezuela, and other elements of the Venezuelan flora are found in a geographically intermediate locality. All three Devonian plant localities in the northwest of South America are within the Colombian Eastern Cordillera and its northern extension, the Venezuelan Perijá Range, an area that has been integrated as a part of the so-called ‘Eastern Andean Terrane’ or ‘Central Andean Province’, supposedly accreted to the autochthonous block of the Guyana Shield during the early Jurassic or before. Although both invertebrates and plants from this terrane have strong affinities to North American and European assemblages, and might be interpreted as implying a Laurussian origin for the Eastern Andean Terrane, the evidence is not yet unequivocal, with some authors postulating an in situ development of this province.

The textures of the chilled marginal gabbro of the Skaergaard intrusion and of the coarser marginal gabbros 25 yards further in are described and related to counts of the number of separate crystals of the chief mineral species per cubic centimetre of the rock. An explanation of the textures and number of crystal individuals is attempted in terms of various degrees of supersaturation in the magma. In the chilled rock abundant nucleation of plagioclase and olivine, presumably at the middle labile stage of supersaturation, is inferred. This is followed by ophitic or poikilitic crystallization of pyroxene and ilmenite about scarce nuclei, produced at lower temperatures, presumably during the early labile stage of supersaturation for these two minerals. The coarser gabbro, further in from the margin, has about half the number of plagioclase crystals per cm.3 compared with the chilled rock and only one twenty-fifth the number of olivine crystals, and these have a tendency to be ophitic in texture. Under these slower cooling conditions, supersaturation apparently still reached into the middle labile region for plagioclase, giving many nuclei, while olivine, like pyroxene, separated under early labile supersaturation conditions and only relatively few nuclei formed. The contrasting textures of the plateau magma-type basalts of Mull and the tholeiitic basalts of Northern Ireland are explained along similar lines.

Three available graphic-correlation analyses are used to calibrate mid-Palaeozoic conodont zonations: Sweet's scheme for the mid- to Upper Ordovician; Kleffner's for the mid- to Upper Silurian; and Murphy & Berry's for the lower and middle Lower Devonian. The scheme of Sweet is scaled by applying the high-precision U-Pb zircon date of Tucker and others for the Rocklandian and linked with that of Kleffner by scaling the graptolite sequence of the Ordovician-Silurian global stratotype section to fit two similarly derived dates from this sequence. The top of Kleffner's scheme, all of Murphy & Berry's, as well as standard zones to the Frasnian are calibrated by using tie-points of the latest Cambridge-BP time-scale (GTS 89). However, the recent microbeam zircon date by Claoué-Long and others for the Hasselbachtal Devonian-Carboniferous auxiliary stratotype is used to calibrate the standard Famennian zones. Also the similarly derived but preliminary determination reported by Roberts and others from the Isismurra Formation of New South Wales is tentatively taken as the top of the Tournaisian and so used to calibrate Tournaisian zones. Despite the considerable extrapolation required to compile these schemes and their inherent errors, the resultant time-scale closely agrees with other dates of Tucker and others from the Llanvirn as well as the GTS 89 Homerian-Gorstian tie-point. This suggests that stratigraphic methods can be usefully applied to geochronometry. The Llandovery appears to have lasted longer (16 m. y.) than usually envisaged and the Ordovician-Silurian boundary may need to be lowered to approximately 443.5 Ma. Certainly, chrons varied widely in duration and further stratigraphic studies to estimate their relative durations as well as high-resolution dating for their calibration will be crucial to more accurate biochronometries.

The links between the Siberian Traps and the end-Permian mass extinction, and between the North Atlantic igneous province (NAIP) and the Paleocene–Eocene thermal maximum (PETM), demonstrate a critical role for large igneous provinces (LIPs) in the disruption of the Earth-surface carbon cycle (ESCC). High-precision ages for both volcanic provinces and the associated environmental crises show that, in both cases, the crisis was contemporaneous with the volcanism. The NAIP comprises two phases: the earlier Phase 1 (c. 61 Ma) and the much more voluminous Phase 2 (c. 56 Ma), linked to the opening of the NE Atlantic. The latter triggered the PETM, the largest Cenozoic hyperthermal. The Siberian Traps are significantly more voluminous than the NAIP, and triggered the end-Permian mass extinction. The masses of volcanic CO2 emitted from these provinces may have been much greater than previously suggested as substantial gas may come from intrusive bodies deep within the crust. Precursory warming due to the accumulation of volcanic CO2 in the atmosphere likely triggered the release of low-δ13C methane hydrate, although the masses of methane hydrate alone may have been insufficient to account for the observed temperature rises. The organic C was likely strongly supplemented by magmatically derived carbon and thermogenic carbon released during emplacement of sills and dykes into C-rich sedimentary units. More data are required on the volcanic flux rates in order to refine the cause–effect relationships between LIPs and the ESCC.

The late Precambrian and Cambrian world experienced explosive evolution of the biosphere, including the development of biomineral skeletons, and notably of phosphate and siliceous skeletons in the initial stages of the adaptive radiation. Ongoing research indicates profound changes in climate and atmospheric carbon dioxide over this span of time. Glacial conditions of the Varangian epoch occur enigmatically at low latitudes, associated with carbonate rocks. Later changes in palaeogeography, sea level rise and salinity stratification encouraged prolonged ‘greenhouse’ conditions in both latest Precambrian and Cambrian times, with indications of relatively low primary production in the oceans. The Precambrian–Cambrian boundary interval punctuated this trend with evaporites, phosphogenic events and carbon isotope excursions; these suggest widespread eutrophication and conjectured removal of carbon dioxide from the atmosphere. Whatever the cause, nutrient–enriched conditions appear to have coincided with the development of phosphatic and siliceous skeletons among the earliest biomineralized invertebrates.

Protector Shoal, the northernmost and most silicic volcano of the South Sandwich arc, erupted dacite–rhyolite pumice in 1962. We report geochemical data for a new suite of samples dredged from the volcano. Geochemically, the dredge and 1962 samples form four distinct magma groups that cannot have been related to each other, and are unlikely to have been related to a single basaltic parent, by fractional crystallization. Instead, the silicic rocks are more likely to have been generated by partial melting of basaltic lower crust within the arc. Trace element and Sr–Nd isotope data indicate that the silicic volcanics have compositions that are more similar to the volcanic arc than the oceanic basement formed at a back-arc spreading centre, and volcanic arc basalts are considered to be the likely source for the silicic magmas. The South Sandwich Islands are one of several intra-oceanic arcs (Tonga–Kermadec, Izu–Bonin) that have: (1) significant amounts of compositionally bimodal mafic–silicic volcanic products and (2) 6.0–6.5 km s−1P-wave velocity layers in their mid-crusts that have been imaged by wide-angle seismic surveys and interpreted as intermediate-silicic plutons. Geochemical and volume considerations indicate that both the silicic volcanics and plutonic layers were generated by partial melting of basaltic arc crust, representing an early stage in the fractionation of oceanic basalt to form continental crust.

The Late Miocene Sorbas Member of the Sorbas Basin, Almería Province, southeast Spain contains an extensive avian ichnofauna preserved in lagoonal marls. Three distinctive avian ichnotaxa can be identified: Antarctichnus fuenzalidae Covacevich and Lamperein, 1970; Iranipeda millumi n. ichnosp.; and Roepichnus grahami n. ichnogen, n. ichnosp. In common with many other Cenozoic avian ichnofaunas, these traces are associated with shorebirds, including plovers, storks, ducks and/or gulls, respectively. Associated mammalian tracks include possible cat and artiodactyl footprints. The avian tracks are abundant and show a range of behavioural aspects in common with other recorded examples of Cretaceous–Recent shorebird tracks. These include both solitary and group activities consistent with their postulated avian tracemakers.

ONE of the objectives of the East African Archaeological Expedition of 1934–5 was the examination of the plateau north of Lake Eyasi—the Vogel River area—where, it was hoped, a continuation of the richly fossiliferous Oldowaya Middle Pleistocene lake beds would be found. This hope was essentially realized ; deposits with an interesting Middle Pleistocene fauna a little earlier than that of Oldoway were found, with the difference that they were entirely terrestrial. The development of precisely datable beds resting upon a main Rift Valley Scarp made possible a study of physiographical changes associated with the faulting.

A newly investigated travertine deposit in southern Tibet provides a window into Holocene hydrological, geomorphic and climatic processes near the boundary of the Tibetan Plateau and High Himalaya. Travertine, deposited as a result of the degassing of CO2-rich groundwater as it emerges on the Earth's surface, is in many cases formed along the trace of major crustal-scale faults in primarily extensional tectonic regimes. A travertine platform measuring roughly 1 km by 0.5 km exists near the town of Nyalam in southern Tibet along a major Himalayan down-to-the-N normal fault, the South Tibetan Fault System. A wide variety of travertine depositional textures and features are recorded in the platform on a series of terraces. Active travertine deposition was observed from spring mounds and seeps along the base of the platform at the modern river level. Palaeotemperatures of spring water, calculated from δ18O of the travertine, range from 9 to 25 °C, which closely matches the temperatures recorded from modern springs in the area. A complex geomorphological landscape records interaction between growing alluvial fans, travertine accumulation, and a rapidly down-cutting river with associated fluvial terraces. River incision was contemporaneous with travertine deposition, as indicated by cemented fluvial river gravel layers interbedded with travertine. High 87Sr/86Sr ratios in the travertine (mean of 0.7168) indicate subsurface fluid interaction with radiogenic crystalline rocks of the underlying Greater Himalaya. Uranium-series ages of the travertine platform range from 5400 a (±950 a) to 11600 a (±1000 a), and indicate a younging progression from higher terraces near the valley wall to lower terraces at present-day river level. Travertine that overlies a river gravel terrace 18 m above river level formed at 11600 a. This date yields a local incision rate of 1.6 mm a−1, consistent with estimated fluvial incision rates in the High Himalaya. The range of our U-series ages coincides with an interval of higher precipitation associated with greater intensity of the Indian monsoon, which led to elevated spring discharge and carbonate precipitation in this part of the High Himalayas.

The Neogene extensional province of southern Tuscany in central Italy provides an outstanding example of fossil and active structurally controlled fluid flow and epithermal ore mineralization associated with post-orogenic silicic magmatism. Characterization of the hydrodynamic regime leading to the genesis of the polysulphide deposit (known as Filone di Boccheggiano) hosted within the damage zone of the Boccheggiano Fault is a key target to assess modes of fossil hydrothermal fluid circulation in the region and, more generally, to provide inferences on fault-controlled hydrothermal fluid flow in extensional settings. We provide a detailed description of the fault zone architecture and alteration/mineralization associated with the Boccheggiano ore deposit and report the results of fluid inclusion and stable oxygen isotope studies. This investigation shows that the Boccheggiano ore consists of an adularia/illite-type epithermal deposit and that sulphide ore deposition was controlled by channelling of hydrothermal fluids of dominantly meteoric origin within the highly anisotropic permeability structure of the Boccheggiano Fault. The low permeability structure of the fault core compartmentalized the fluid outflow preventing substantial cross-fault flow, with focused fluid flow occurring at the hangingwall of the fault controlled by fracture permeability. Fluid inclusion characteristics indicate that ore minerals were deposited between 280° and 350°C in the upper levels of the brittle extending crust (lithostatic pressure in the order of 0.1 GPa). Abundant vapour-rich inclusions in ore-stage quartz are consistent with fluid immiscibility and boiling, and quartz ore vein textures suggest that mineralization in the Boccheggiano ore deposit occurred during cyclic fluid flow in a deformation regime regulated by transient and fluctuating fluid pressure conditions. Results from this study (i) predict a strongly anisotropic permeability structure of the fault damage zone during crustal extension, and (ii) indicate the rate of secondary (structural) permeability creation and maintenance by active deformation in the hangingwall of extensional faults as the major factor leading to effective hydraulic transmissivity in extensional terranes. These features intimately link ore-grade mineralization in extensional settings to telescoping of hydrothermal flow along the hangingwall block(s) of major extensional fault zones.

Arrested-type charnockite formation occurs in an assemblage of high-grade gneisses at several localities of the Chilka Lake area that belongs to the Proterozoic Eastern Ghats Belt of India. The isolated ellipsoidal domains are found exclusively in leucogranite (leptynite) bands that intruded lit-par- lit interbanded granulite-grade supracrustal and intermediate igneous rocks (khondalite–enderbite). Macrostructures and microfabrics document a multiple deformation of the rock assemblage under high-grade conditions. The intrusion of the leucogranitic melts separates a first episode of deformation, D1, from a younger progressive deformation, D2–D4. A transpressive regime and inhomogeneous deformation is indicated for D2–D4 by the associated structures and fabrics. But quartz c-axis patterns show that pure shear prevailed during the closing stages of deformation. The spatial distribution and orientation of the ellipsoidal charnockite domains within the host leptynite and the orientation pattern of orthopyroxene c-axes inside the domains provide evidence for a synkinematic in situ formation of the domains during D3, through partial breakdown of the leptynite assemblage (Bt+Grt+Qtz+Fl1[rlhar ]Opx+Fsp+Ilm+Fl2/L). Local fluid migration along steep foliation planes associated with large-scale D3 folds triggered the reaction. Orthopyroxene blastesis was confined to the centre of the domains, and an envelope formed in which the residing fluid caused secondary intergranular formation of chlorite, ore and carbonate, imparting the domains' typical greenish-brown charnockite colour. The shape of the envelope, which varies from prolate in limbs to oblate in hinges of D3 folds, is responsive to the local stress field. Comparison of chemical rock compositions supports the in situ formation of charnockite in leptynite. Subtle compositional differences are controlled by the changing mineralogy. Compared to the host leptynite, the charnockite domains are enriched in K2O, Ba, Rb and Sr, but depleted in FeO*, MnO, Y and Zr. The data obtained in this study provide conclusive evidence that the ellipsoidal charnockite domains do not represent remnants of stretched enderbite layers as proposed by Bhattacharya, Sen & Acharyya, but formed in situ in the leptynite as a result of localized synkinematic fluid migration late in the deformation history.

New data implying crustal activation of Eastern Avalonia along the Anglo-Brabant fold belt are presented. Late Ordovician subduction-related magmatism in East Anglia and the Brabant Massif, coupled with accelerated subsidence in the Anglia Basin and in the Brabant Massif during Silurian time, indicate a foreland basin development. Final collision resulted in folding, cleavage development and thrusting during the mid-Lochkovian to mid-Eifelian. In the southeast of the Anglo-Brabant fold belt, Acadian deformation produced basin inversion and the regional antiformal structure of the Brabant Massif. The uplift, inferred from the sedimentology, petrography and reworked palynomorphs in the Lower Devonian of the Dinant Synclinorium is confirmed by illite crystallinity studies. The tectonic model discussed implies the presence of two subduction zones in the eastern part of Eastern Avalonia, one along the Anglo-Brabant fold belt and another under the North Sea in the prolongation of the North German–Polish Caledonides.

Detailed dinoflagellate cyst analysis of the Lower–Middle Miocene Berchem Formation at the southernmost margin of the North Sea Basin (northern Belgium) allowed a precise biostratigraphical positioning and a reconstruction of the depositional history. The two lower members of the formation (Edegem Sands and decalcified Kiel Sands) are biostratigraphically regarded as one unit since no significant break within the dinocyst assemblages is observed. The base of this late (or latest) Aquitanian–Burdigalian unit coincides with sequence boundary Aq3/Bur1 as defined by Hardenbol and others, in work published in 1998. A hiatus at the Lower–Middle Miocene transition separates the upper member (the Antwerpen Sands) from the underlying member. The greater part of the Antwerpen Sands were deposited in a Langhian (latest Burdigalian?)–middle Serravallian interval. The base of this unit coincides with sequence boundary Bur5/Lan1. Biostratigraphical correlation points to a diachronous post-depositional decalcification within the formation since parts of the decalcified Kiel Sands can be correlated with parts of the calcareous fossil-bearing section, up to now interpreted as Antwerpen Sands. The dinoflagellate cyst assemblages are dominated by species with a inner neritic preference, although higher numbers of oceanic taxa in the upper part of the formation indicate incursions of oceanic watermasses into the confined depositional environment of the southern North Sea Basin.

The terrestrialization of life has profoundly affected the biosphere, geosphere and atmosphere, and the Geological Magazine has published key works charting the development of our understanding of this process. Integral to this understanding – and featuring in one of the Geological Magazine publications – is the Devonian Rhynie chert Konservat-Lagerstätte located in Aberdeenshire, Scotland. Here we provide a review of the work on this important early terrestrial deposit to date. We begin by highlighting contributions of note in the Geological Magazine improving understanding of terrestrialization and Palaeozoic terrestrial ecosystems. We then introduce the Rhynie chert. The review highlights its geological setting: the Caledonian context of the Rhynie Basin and its nature at the time of deposition of the cherts which host its famous fossils. There follows an introduction to the development of the half-graben in which the cherts and host sediments were deposited, the palaeoenvironment this represented and the taphonomy of the fossils themselves. We subsequently provide an overview of the mineralization and geochemistry of the deposit, and then the fossils found within the Rhynie chert. These include: six plant genera, which continue to provide significant insights into the evolution of life on land; a range of different fungi, with recent work starting to probe plant–fungus interactions; lichens, amoebae and a range of unicellular eukaryotes and prokaryotes (algae and cyanobacteria); and finally a range of both aquatic and terrestrial arthropods. Through continued study coupled with methodological advances, Rhynie fossils will continue to provide unique insights into early life on land.

In March of the present year (1909) the Rev. R. Ashington Bullen, who was then travelling in Majorca, wrote to inform me of the existence of a supposed ossiferous breccia situated on the east coast of that island. This caused a long-deferred project of exploring the Balearic Islands in search of Pleistocene cave-deposits to be carried out, and a month was spent in this manner, although a much longer period would be required for an exhaustive examination of Majorca alone, which was the only one of the group visited. A considerable tract of limestone country was investigated, and the result was the discovery of three ossiferous cave-deposits, including the one mentioned above. Similar remains were procured from each of these, and on examination prove to be those of an ungulate which appears to be without parallel. No great quantity of material was obtained, but this fortunately includes a well-preserved skull of an old individual with the associated mandible and atlas vertebra—the typespecimen, see Fig. 1—as well as a number of perfect examples of various limb-bones.