Hostname: page-component-cd9895bd7-8ctnn Total loading time: 0 Render date: 2024-12-26T05:05:57.914Z Has data issue: false hasContentIssue false

Crinoids from the Lower (Pragian–Emsian) and Middle (early Eifelian) Devonian of Bolivia (Icla and Belén formations, Malvinokaffric Realm)

Published online by Cambridge University Press:  21 September 2020

Sandro M. Scheffler*
Affiliation:
Laboratório de Paleoinvertebrados – LAPIN, Departamento de Geologia e Paleontologia, Museu Nacional, Universidade Federal do Rio de Janeiro - UFRJ, Horto do Museu Nacional, Parque Quinta da Boa Vista, CEP 20.940-040, São Cristóvão, Rio de Janeiro, RJ, Brazil
*
*Corresponding Author.

Abstract

Two new families and three new genera and species of Crinoidea from the Icla and Belén formations, Bolivian Devonian, are described. The material is identified as Meperocrinus angelina n. gen. n. sp., from the Icla Formation of the Sub-Andina region, Tuberocrinus lapazensis n. gen. n. sp. and Aenigmaticumcrinus rochacamposi n. gen. n. sp., both from the Belén Formation of the Bolivian Altiplano. Meperocrinus angelina n. gen. n. sp. is an Emperocrinidae, a family described from the Silurian of the United States of America. Tuberocrinus lapazensis n. gen. n. sp. is assigned to a new family of Dimerocrinitacea, the Tuberocrinidae n. fam., which is closely related to Pterinocrinidae that already has representatives in Bolivia (Apurocrinus sucrei McIntosh), Argentina (Pterinocrinus? australis Haude), and Colombia (Bogotacrinus scheibei McIntosh). Aenigmaticumcrinus rochacamposi n. gen. n. sp. is also as assigned to a new family of Dimerocrinitacea, the Aenigmaticumcrinidae n. fam.; a family with very distinctive features. These data demonstrate that the Malvinokaffric crinoid fauna in Bolivia has a mixed origin, with forms that arrived in South America from Europe (Old World Realm) and United States of America (Eastern America Realm) during the late Silurian or earliest Devonian, as well as a local stock, which evolved to highly endemic lineages.

UUID: http://zoobank.org/f03120c6-264a-4ae9-81e0-46bfe844e65e.

Type
Articles
Copyright
Copyright © The Author(s), 2020. Published by Cambridge University Press on behalf of The Paleontological Society

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Ausich, W.I., 1987, Revision of Rowle's Ordovician(?) and Silurian crinoids from Missouri: Journal of Paleonotology, v. 6, p. 563578.10.1017/S0022336000028729CrossRefGoogle Scholar
Ausich, W.I., 1998, Phylogeny of Arenig to Caradoc crinoids (Phylum Echinodermata) and suprageneric classification of the Crinoidea: The University of Kansas Paleontological Contributions, no. 9, 36 p.Google Scholar
Ausich, W.I., 2009, These are not the crinoids your daddy knew!, in Catalani, J., and Cozart, C., eds., Crinoids: Mid-America Paleontology Society Digest, v. 32, p. 622.Google Scholar
Ausich, W.I., and Dravage, P., 1988, Crinoids from the Brassfield Formation of Adams County, Ohio: Journal of Paleontology, v. 62, p. 285289.10.1017/S0022336000029930CrossRefGoogle Scholar
Ausich, W.I., Kammer, T.W., and Baumiller, T.K., 1994, Demise of the middle Paleozoic crinoid fauna: a single extinction event or rapid faunal turnover?: Paleobiology, v. 20, p. 345361.10.1017/S0094837300012811CrossRefGoogle Scholar
Baumiller, T.K., 1994, Patterns of dominance and extinction in the record of Paleozoic crinoids, in David, B., Guille, A., Féral, J.P., and Roux, M., eds., Echinoderms Through Time: Rotterdam, A.A. Balkema, p. 193198.Google Scholar
Baumiller, T.K., and Plotnick, R.E., 1989, Rotational stability in stalked crinoids and the function of wing plates in Pterotocrinus depressus: Lethaia, v. 22, p. 317326.10.1111/j.1502-3931.1989.tb01348.xCrossRefGoogle Scholar
Blieck, A.P., Gagnier, Y., Bigey, F.P., Edgecombe, G.D., Janvier, P., Loboziak, S., Racheboeuf, P.R., Sempere, T., and Steemans., P., 1996, New Devonian fossil localities in Bolivia: Journal of South American Earth Sciences, v. 9, p. 295308.10.1016/S0895-9811(96)00015-6CrossRefGoogle Scholar
Boucot, A.J., 1971, Malvinokaffric Devonian marine community: distribution and implications for Godwana: 1° Simpósio Brasileiro de Paleontologia, 1970, Anais da Academia Brasileira de Ciências, v. 43 (suplemento), p. 2349.Google Scholar
Boucot, A.J., 1988, Devonian biogeography: an update, in Mcmillan, N.J., Embry, A.F., and Glass, D.J., eds., Devonian in the World: Canadian Society of Petroleum Geologists, Memoir 14, p. 211227.Google Scholar
Boucot, A.J., and Racheboeuf, P.R., 1993, Biogeographic summary of the Malvinokaffric Realm Silurian and Devonian fossils, in Suarez-Soruco, R., ed., Fosiles y Facies de Bolivia: Revista Técnica de YPFB, II Invertebrados y Paleobotanica, v. 13–14, no. 1–4, p. 7175.Google Scholar
Boucot, A.J., Johnson, J.G., and Talent, J.A., 1969, Early Devonian brachiopod zoogeography: The Geological Society of America, Special Papers, no. 119, 114 p.10.1130/SPE119-p1CrossRefGoogle Scholar
Branisa, L., 1965, Los fosiles guias de Bolivia: Paleozoic: Boletin Servicio Geologico de Bolivia, v. 6, p. 1282.Google Scholar
Breimer, A., 1962, A monograph on Spanish Paleozoic crinoidea: Overdruk uit Leidse Geologische Mededelingen, Deel 27, p. 1190.Google Scholar
Breimer, A., 1978, General morphology: recent crinoids, in Moore, R.C., and Teichert, C., eds., Treatise on Invertebrate Paleontology. Part T. Echinodermata 2, v. 1. Lawrence, The University of Kansas Printing Service, p. T9T58.Google Scholar
Breimer, A., and Macurda, D.B. Jr., 1972, The Phylogeny of Fissiculate Blastoids: Verhangelingen der Koninklijke Nederlandsche Akademie van Wetenschappen, Afdeling Natuurkunde, Eerste Reeks, Deel 26, 390 p.Google Scholar
Brett, C.E., 1978, Description and paleoecology of a new lower Silurian camerate crinoid: Journal of Paleontology, v. 5, p. 91103.Google Scholar
Brower, J.C. 1994, Camerate crinoids from the Middle Ordovician (Galena Group, Dunleith Formation) of northern Iowa and southern Minnesota: Journal of Paleontology, v. 68, p. 570599.10.1017/S0022336000025932CrossRefGoogle Scholar
Carvalho, M.G.P., and Ponciano, L.C.M.O., 2015, The Devonian trilobites of Brazil: a summary: Journal of South American Earth Sciences, v. 64, p. 217228.10.1016/j.jsames.2015.10.010CrossRefGoogle Scholar
Carvalho, M.GP., Edgecombe, G.D., and Smith, L., 2003, New calmoniid trilobites (Phacopina: Acastoidea) from the Devonian of Bolivia: American Museum Novitates, no. 3407, p. 117.10.1206/0003-0082(2003)407<0001:NCTPAF>2.0.CO;22.0.CO;2>CrossRefGoogle Scholar
Clarke, J.M., 1913, Fósseis Devonianos do Paraná: Serviço Geológico e Mineralógico do Brasil, Monografia 1, 353 p.Google Scholar
Cole, S.R., 2017, Phylogeny and morphologic evolution of the Ordovician Camerata (Class Crinoidea, Phylum Echinodermata): Journal of Paleontology, v. 91, p. 815828.10.1017/jpa.2016.137CrossRefGoogle Scholar
Di Pasquo, M., Amenábar, C.R., and Noetinger, S., 2009, Middle Devonian microfloras and megafloras from western Argentina and southern Bolivia: their importance in the palaeobiogeographical and palaeoclimatic evolution of western Gondwana, in Konigshof, P., ed., Devonian Change: Case Studies in Palaeogeography and Palaeoecology: The Geological Society, London, Special Publications, v. 314, p. 193213.Google Scholar
Eckert, J.D., 1984, Early Llandovery crinoids and stelleroids from the Cataract Group (lower Silurian), southern Ontario, Canada: Royal Ontario Museum Life Sciences, Contributions, no. 137, p. 183.Google Scholar
Eckert, J.D., and Brett, C., 2001, Early Silurian (Llandovery) Crinoids from the Lower Clinton Group, Western New York State: Bulletins of American Paleontology, no. 360, 88 p.Google Scholar
Eldredge, N., and Branisa, L., 1980, Calmoniid trilobites of the Lower Devonian Scaphiocoelia Zone of Bolivia, with remarks on related species: Bulletin of the American Museum of Natural History, v. 165, p. 181289.Google Scholar
Fonseca, V.M.M., 2004, Chonetoidea (Brachiopoda) do Devoniano Médio das bacias do Amazonas e Parnaíba, Brasil: Arquivos do Museu Nacional, v. 62, p. 193215.Google Scholar
Fonseca, V.M.M., and Melo, J.H.G., 1987, Ocorrência de Tropidoleptus carinatus (Conrad) (Brachiopoda, Orthida) na Formação Pimenteira, e sua importância paleobiogeográfica, 10th Congresso Brasileiro de Paleontologia, Anais, v. 2, p. 505537.Google Scholar
Frest, T.J., and Strimple, H.L., 1981, New camerate crinoids from the Silurian of North America: Journal of Paleontology, v. 55, p. 639655.Google Scholar
Grahn, Y., 2002, Upper Silurian and Devonian Chitinozoa from central and southern Bolivia, central Andes: Journal of South American Earth Sciences, v. 15, p. 315326.10.1016/S0895-9811(02)00045-7CrossRefGoogle Scholar
Grahn, Y., and Gutiérrez, P.R., 2001, Silurian and Middle Devonian Chitinozoa from the Zapla and Santa Bárbara ranges, Tarija Basin, Northwest Argentina: Ameghiniana, v. 38, p. 3550.Google Scholar
Haude, R., 1995, Echinodermen aus dem Unter-Devon der arentinischen Präkordillere: Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, v. 197, no. 1, p. 3786.10.1127/njgpa/197/1995/37CrossRefGoogle Scholar
Haude, R., 2004, Morphology and paleobiology of echinoderms in the Lower Devonian of the Argentine Precordillera, in Heinzeller, T., and Nebelsick, J.H., eds., Echinoderms: London, Francis and Taylor Group, p. 417420.Google Scholar
Isaacson, P.E., 1977, Devonian stratigraphy and brachiopod paleontology of Bolivia, Part A. Orthida and Strophomenida: Palaeontographica Abteilung A, Palaeozoologie-Stratigraphie, v. 155, p. 133192.Google Scholar
Isaacson, P.E., and Perry, D.G., 1977, Biogeography and morphological conservatism of Tropidoleptus (Brachiopoda, Orthida) during the the Devonian: Journal of Paleontology, v. 51, p. 11081122.Google Scholar
Isaacson, P.E., and Sablock, P.E., 1990, Devonian paleogeography and palaeobiogeography of the Central Andes: Geological Society of London, Memoir, v. 12, p. 431435.10.1144/GSL.MEM.1990.012.01.40CrossRefGoogle Scholar
Jaekel, O., 1895, Beiträge zur Kenntniss der palaeozoischen Crinoiden Deutschlands: Paläontologisches Abhandlungen, v. 7, n.f. 3, p. 1116.Google Scholar
Jaekel, O., 1918, Phylogenie und System der Pelmatozoen: Paläeontologische Zeitschrift, v. 3, p. 1128.Google Scholar
Jell., P.A., 1999, Silurian and Devonian crinoids from central Victoria: Memoirs of' the Queensland Museum v. 43, p. 1114.Google Scholar
Jell, P.A., and Theron, J.N., 1999, Early Devonian echinoderms from South Africa: Memoirs of the Queensland Museum, v. 43, p. 115199.Google Scholar
Katzer, F., 1897, A fauna devonica do Rio Maecuru, e suas relacões com a fauna de outros terrenos devonicos do globo: Boletim do Museu Paraense de História Natural e Ethnographia, v. 11, p. 204236.Google Scholar
Katzer, F., 1903, Grundzuge der Geologie des unteren Amazonasgebietes (des States Pará in Brasilien). Leipzig, Max Weg, 298 p.Google Scholar
Katzer, F., 1933, Geologia do Estado do Pará: Boletim do Museu Paraense Emílio Goeldi de História Natural e Etnografia, v. 9, p. 1269. [Tradução frei Hugo Mense, do original alemão: Katzer, F., 1903. Grundzüge der Geologie Unteren Amazonasgebietes.]Google Scholar
Kirk, E., 1913, Crinoidea, in Clarke, J.M., ed., Fósseis Devonianos do Paraná: Serviço Geológico e Mineralógico do Brasil, Monografia 1, p. 316318.Google Scholar
Knod, R., 1908, Devonische Faunen Boliviens: Neues Jahrbuch für Mineralogie, Geologie und Paäontologie, v. 25, p. 493600.Google Scholar
Leme, J.M., Simões, M.G., and Van Iten, H., 2006, Reinterpretation of Malvinoconularia cahuanotensis (Braniša and Vanek) from the Devonian Altiplano and Western Andean Cordillera, Bolivia, South America: Ameghiniana, v. 43, p. 285292.Google Scholar
Le Menn, J., 1980, Les Crinoides, in Plusquellec, Y., ed., Les Schistes et calcaires de l'Armorique: Devonien inferieur, Massif armoricain: sedimentologie, paleontologie, stratigraphie: Memoires de la Societe geologique et mineralogique de Bretagne, v. 23, p. 249271.Google Scholar
Le Menn, J., 1990, Présence de Monstrocrinus (Crinoidea) dans l' Emsian Supérieur du domaine Nord-Gondwanien (Espagne, Algerie): Annales de Paléontologie, v. 76, p. 149158.Google Scholar
Lyon, S.S., and Casseday, S.A., 1860, Description of nine new species of Crinoidea from the Subcarboniferous rocks of Indiana and Kentucky: American Journal Science, ser. 2, no. 29, p. 6879.10.2475/ajs.s2-29.85.68CrossRefGoogle Scholar
Macurda, D.B., 1979, The Devonian blastoids of Bolivia: Journal of Paleontology, v. 53, p. 13611373.Google Scholar
Mcintosh, G.C., 1981, Apurocrinus sucrei, a new genus and species of camerate crinoid from the Lower Devonian of Bolivia: Journal of Paleontology, v. 55, p. 948952.Google Scholar
McIntosh, G.C., 1987, Review of the Devonian camerate crinoid Bogotacrinus scheibei Schimdt from Colombia: Journal of Paleontology, v. 61, p. 750757.10.1017/S0022336000029103CrossRefGoogle Scholar
McIntosh, G. C., 1988, Boliviacrinus isaacsoni, a new genus and species of Middle Devonian camerate crinoid from Bolivia: Journal of Paleontology, v. 62, p. 622626.Google Scholar
Melo, J.H.G. de, 1985, A Província Malvinocáfrica no Devoniano do Brasil [M.Sc. thesis: estado atual dos conhecimentos]: Rio de Janeiro, Universidade Federal do Rio de Janeiro, Instituto de Geociências, 890 p.Google Scholar
Melo, J.H.G. de., 1988, The Malvinokaffric Realm in the Devonian of Brazil, in Mcmillan, N.J., Embry, A.F., and Glass, O.J., eds., Devonian of the World: Canadian Society of Petroleum Geologists, Memoir 14, p. 667703.Google Scholar
Miller, J.S., 1821, A Natural History of the Crinoidea, or lily-shaped animals, with observations of the genera Asteria, Euryale, Comatula and Marsupites: Bristol, C. Frost, 50 p.10.5962/bhl.title.32130CrossRefGoogle Scholar
Miller, S.A., and Gurley, W.F.E., 1895, Description of new species of Palaeozoic Echinodermata: Illinois State Museum, Bulletin 6, p. 162.10.5962/bhl.title.119753CrossRefGoogle Scholar
Moore, R.C., and Laudon, L.R., 1943, Evolution and classification of Paleozoic crinoids: Geological Society of America, Special Papers, v. 46, 153 p.10.1130/SPE46-p1CrossRefGoogle Scholar
Morzadec, P., Pans, F., Plusquellec, Y., Racheboeuf, P.R., and Weyant, M., 1988, Devonian stratigraphy and paleogeography of the Armorican Massif (N.W. France), in Mcmillan, N.J., Embry, A.F., and Glass, D.J., eds., Devonian of the World: Canadian Society of Petroleum Geologists, Memoir 14, p. 401420.Google Scholar
Plusquellec, Y., Boumendjel, K., Morzadec, P., and Paris., F., 1997, Les Faunes Dévoniennes d'Ougarta dans la paléogéographie des regions Maghrébo-Européennes: Annales de Societé Géologique du Nord, v. 5, p. 123128.Google Scholar
Prokop, R.J., 2010, Aishacrinidae, a new family of camerate crinoids from the Silurian of Bohemia, Czech Republic: Journal of the National Museum, Natural History Series, v. 179, p. 4146.Google Scholar
Prokop, R.J., and Petr, V., 1989, New Silurian and Devonian crinoid taxa from Bohemia: Časopis pro Mineralogii a Geologii, v. 34, p. 4552.Google Scholar
Racheboeuf, P.R., Le Hérissé, A., Paris, F., Babin, C., Guillocheau, F., Truyols-Massoni, M., and Suarez-Soruco, R., 1993, Le Devonien de Bolivie: biostratigraphie et chronostratigraphie: Comptes-Rendus de l'Académie des Sciences, Paris, v. 317, p. 795802.Google Scholar
Robardet, M., 2003, The Armorica ‘microplate’: fact or fiction? Critical review of the concept and contradictory palaeobiogeographical data: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 195, p. 125148.10.1016/S0031-0182(03)00305-5CrossRefGoogle Scholar
Robardet, M., Blaise, J., Bouyx, E., Gourvennec, R., Lardeaux, H., Le Herrisse, A., Le Menn, J., Melou, M., Paris, F., Plusquellec, Y., Poncet, J., Regnault, S., Rioult, M., and Weyant, M., 1993, Paléogéographie de l'Europe occidentale de l'Ordovicien au Devonian: Bulletin de la Societé géologique de France, v. 164, p. 683695.Google Scholar
Roemer, C.F. 1855, Palaeozoic Pelmatozoa: Lethaea Geognostica, v. 2, p. 180.Google Scholar
Rowley, R.R., 1904, The Echinodermata of the Missouri Silurian and a new brachiopod: American Geologist, v. 34, p. 269282.Google Scholar
Salter, J.W., 1856, Description of Paleozoic Crustacea and Radiata from South Africa: Transactions of the Geological Society of London, v. 7, p. 215224.Google Scholar
Scheffler, S.M., 2007, Um novo crinoide do Devoniano Inferior da Bolívia (Formação Icla): Anais do 20° Congresso Brasileiro de Paleontologia, Búzios, p. 297.Google Scholar
Scheffler, S.M., 2015, Stalked echinoderms of the Brazilian Devonian and their palaeobiogeographical affinities: Boletim do Museu Paraense Emílio Goeldi, Ciências Naturais, v. 10, p. 6381.Google Scholar
Scheffler, S.M., and Fernandes, A.C.S., 2007a, Crinoidea da Formação Ponta Grossa (Devoniano, Bacia do Paraná), Brasil: Arquivos do Museu Nacional, v. 65, p. 8398.Google Scholar
Scheffler, S.M., and Fernandes, A.C.S., 2007b, Blastoidea da Formação Ponta Grossa (Devoniano, Bacia do Paraná), Brasil: Arquivos do Museu Nacional, v. 65, p. 99112.Google Scholar
Scheffler, S.M., Fernandes, A.C.S., and Fonseca, V.M.M., 2006, Crinoidea da Formação Maecuru (Devoniano da Bacia do Amazonas), Estado do Pará, Brasil: Revista Brasileira de Paleontologia, v. 9, p. 235242.10.4072/rbp.2006.2.07CrossRefGoogle Scholar
Scheffler, S.M., Silva, S.D., Gama, J.M. Jr., Fonseca, V.M.M., and Fernandes, A.C.S., 2011, Middle Devonian crinoids from the Parnaiba Basin (Pimenteira Formation, Tocantins State, Brazil): Journal of Paleontology, v. 85, p. 11881198.10.1666/10-142.1CrossRefGoogle Scholar
Scheffler, S.M., Fernandes, A.C.S., and Fonseca, V.M.M., 2013, Alguns Crinoides da Formação Ponta Grossa e suas afinidades paleobiogeográficas (Devoniano Inferior, Bacia do Paraná, Brasil): Terr@Plural, Ponta Grossa, v. 7, Número Especial, p. 85114.10.5212/TerraPlural.v.7iEspecial.0007CrossRefGoogle Scholar
Scheffler, S.M., Fernandes, A.C.S., and Fonseca, V.M.M., 2014, Crinoids columnals (Echinodermata) of the Ererê Formation (late Eifelian–early Givetian, Amazon Basin), State of Pará, Brazil: Journal of South American Earth Sciences, v. 49, p. 6372.10.1016/j.jsames.2013.11.001CrossRefGoogle Scholar
Scheffler, S.M., Fonseca, V.M.M., and Fernandes, A.C.S., 2015. New crinoids from the Maecuru Formation (middle Eifelian, Amazon Basin), state of Pará, Brazil: Geobios, v. 48, p. 5769.10.1016/j.geobios.2014.11.002CrossRefGoogle Scholar
Távora, V.A., Dias, J.J., and Ramalho, L.V., 2017, Considerações Paleobiogeográficas dos Corais e Briozoários do Devoniano Brasileiro: Anuário do Instituo de Geociências, v. 40, p. 226234.10.11137/2017_1_226_234CrossRefGoogle Scholar
Thompson, J.R, Ausich, W.I., and Smith, L., 2013, Echinoderms from the Lower Devonian (Emsian) of Bolivia (Malvinokaffric Realm): Journal of Paleontology, v. 87, p. 166175.10.1666/12-068R.1CrossRefGoogle Scholar
Thomson, C.W., 1864, Sea lilies: The Intellectual Observer, v. 6, p. 111.Google Scholar
Tobenski, T.L., 2012, Pterotocrinus of the Menard Limestone and Kinkaid Formation (Elviran Stage) Chesterian Series in the Illinois Basin [M.Sc. Thesis]: Carbondale, Illinois, Southern Illinois University, Department of Geology, 104 p.Google Scholar
Troth, I., Marshall, J.E.A., Racey, A., and Becker, R.T., 2011, Devonian sea-level change in Bolivia: a high palaeolatitude biostratigraphical calibration of the global sea-level curve: Palaeogeography, Palaeoclimatology, Palaeoecology v. 304, p. 320.10.1016/j.palaeo.2010.10.008CrossRefGoogle Scholar
Ubaghs, G., 1978, Camerata, in Moore, R.C., and Teichert, C., eds., Treatise on Invertebrate Paleontology. Part. T, Echinodermata 2: Lawrence, Kansas, Geological Society of America and University of Kansas Press, p. T408T519.Google Scholar
Von Zittel, K.A., 1876–80, Handbuch der Palaeontologie, v. 1, Palaeozoologie: München, Leipzig, R. Oldenbourg, (1879), no. 1, p. 1765.Google Scholar
Wachsmuth, C., and Springer, F., 1885, Discussion of the classification and relations of the brachiate crinoids, and conclusion of the generic descriptions: Proceedings of the Academy of Natural Sciences of Philadelphia, pt. III, sec. 1, p. 225364. [separate re-paged, 1–138.]Google Scholar
Waisfeld, B.G., 1989, Comunidad de Bridgerocrinus (Crinoidea: Scytalocrinidae) en la Formacion Talàcasto (Devonico) de la precordillera de San Juan, Argentina: Ameghiniana, v. 25, p. 273279.Google Scholar
Webster, G.D., 2000, Palaeobiogeography of Devonian and Carboniferous crinoids faunas of Gondwana: Records of the Western Australian Museum, v. 58 (suppl.), p. 403420.Google Scholar
Welch, J.R., 1978, Pterotocrinus rugosus Lyon and Casseday from the middle Chesterian of the Illinois Basin: Journal of Paleontology, v. 52, p. 904915.Google Scholar
Weller, S., 1900, The paleontology of the Niagaran Limestone, in the Chicago area, the Crinoidea, Part 1 of the Natural History Survey: Chicago Academy of Sciences, Bulletin, v. 4, p. 1153.Google Scholar
Wright, D.F., Ausich, W.I., Cole, S.R., Peter, M.E., and Rhenberg, E.C., 2017, Phylogenetic taxonomy and classification of the Crinoidea (Echinodermata): Journal of Paleontology, v. 91, p. 829846.10.1017/jpa.2016.142CrossRefGoogle Scholar