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Disparid and hybocrinid crinoids (Echinodermata) from the Upper Ordovician (lower Katian) Brechin Lagerstätte of Ontario

Published online by Cambridge University Press:  18 June 2018

William I. Ausich ,
David F. Wright ,
Selina R. Cole  and
Joseph M. Koniecki
William I. Ausich
Affiliation:
School of Earth Sciences, 125 South Oval Mall, The Ohio State University, Columbus, Ohio 43210 〈ausich.1@osu.edu〉
David F. Wright
Affiliation:
Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, P.O. Box 37012, MRC 121, Washington, DC 20013-7012 〈wrightda@si.edu〉 〈colesel@si.edu〉
Selina R. Cole
Affiliation:
Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, P.O. Box 37012, MRC 121, Washington, DC 20013-7012 〈wrightda@si.edu〉 〈colesel@si.edu〉
Joseph M. Koniecki
Affiliation:
3529 E. Joy Rd, Ann Arbor, Michigan 48105 〈paleojk@gmail.com〉
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Abstract

The Brechin Lagerstätte (Katian, Ordovician) from the Lake Simcoe region of Ontario, Canada contains a diverse array of echinoderms. Here, we describe seven disparid and two hybocrinid crinoids (subclass Pentacrinoidea, infraclass Inadunata), including a new disparid species belonging to the Anomalocrinidae (order Homocrinida). In total, the disparids include Anomalocrinus astrictus n. sp.; Cremacrinus guttenbergensis Kolata, 1975; C. inaequalis Billings, 1859; Daedalocrinus bellevillensis Billings, 1883; Eustenocrinus springeri Ulrich, 1925; Iocrinus trentonensis Walcott, 1883; and Isotomocrinus tenuis Billings, 1857b. The hybocrinids include Hybocrinus tumidus Billings, 1857a and Hybocystites problematicus Wetherby, 1880. Previously known from only the holotype, three additional specimens of E. springeri expand our understanding of this unusual crinoid. Nomenclatural acts include: (1) the recommended designation of D. kirki Ulrich, 1925 as a junior synonym of D. bellevillensis is followed; (2) Hybocrinus pristinus Billings, 1858 is designated as a junior synonym of H. tumidus, and previous decisions are followed to retain Hybocystites eldonensis (Parks, 1908) as a junior synonym of H. problematicus; (3) although probably assignable to Anomalocrinus Meek and Worthen, 1865, the aberrant crinoid Glaucocrinus falconeri Parks and Alcock, 1912, and its genus Glaucocrinus Parks and Alcock, 1912, are designated as nomena dubia; (4) Iocrinus similis (Billings, 1857) is also designated as a nomen dubium; and (5) Iocrinus subcrassus torontoensis Fritz, 1925 is designated a junior synonym of I. subcrassus Meek and Worthen, 1865.

UUID: http://zoobank.org/90f8580b-ed7e-4405-97b2-73c9069de7f5

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Articles
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Journal of Paleontology , Volume 92 , Issue 5 , September 2018 , pp. 850 - 871
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Copyright © 2018, The Paleontological Society 

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References

Armstrong, D.K., 2000, Paleozoic geology of the northern Lake Simcoe area, south-central Ontario: Ontario Geological Survey, Open File Report 6011, 34 p.Google Scholar
Arendt, Y.A., 1981, Trekhrukie morskie lilii [Three-armed crinoids]: Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta, no. 189, 195 p.Google Scholar
Ausich, W.I., 1996, Crinoid plate circlet homologies: Journal of Paleontology, v. 70, p. 955–964.Google Scholar
Ausich, W.I., 1998, Phylogeny of Arenig to Caradoc crinoids (phylum Echinodermata) and suprageneric classification of the Crinoidea: University of Kansas Paleontological Contributions, n. ser., v. 9, 36 p.Google Scholar
Ausich, W.I., and Copper, P., 2010, The Crinoidea of Anticosti Island, Québec (Late Ordovician to Early Silurian): Palaeontographic Canadiana, no. 29, 157 p.Google Scholar
Ausich, W.I., and Deline, B., 2012, Macroevolutionary transition in crinoids following the Late Ordovician extinction event (Ordovician to Early Silurian): Palaeogeography, Palaeoclimatology, Palaeoecology, v. 361, p. 38–48. doi: 10.1016/j.palaeo.2012.07.022.Google Scholar
Ausich, W.I., Bolton, T.E., and Cummings, L.M., 1998, Whiterockian (Ordovician) crinoid fauna from the Table Head Group, western Newfoundland, Canada: Canadian Journal of Earth Science, v. 35, p. 121–130.Google Scholar
Ausich, W.I., Brett, C.E., Hess, H., and Simms, M.J., 1999, Crinoid form and function, in Hess, H., Ausich, W.I., Brett, C.E., and Simms, M.J. Fossil Crinoids: Cambridge, UK, Cambridge University Press, p. 3–30.Google Scholar
Ausich, W.I., Kammer, T.W., Rhenberg, E.C., and Wright, D.F., 2015, Early phylogeny of crinoids within the Pelmatozoan clade: Palaeontology, v. 58, p. 937–952. doi: 10.1111/pala.12204.Google Scholar
Bassler, R.S., 1915, Bibliographic index of American Ordovician and Silurian fossils: United States National Museum Bulletin, v. 92, pt. 1, p. 1–718; pt. 2, p. 719–1521.Google Scholar
Bassler, R.S., 1938, Pelmatozoa Palaeozoica, in Quenstedt, W., ed., Fossilium Catalogus, I, Animalia, Part 83: ‘s-Gravenhage: The Netherlands, W. Junk, 194 p.Google Scholar
Bassler, R.S., and Moodey, M.W., 1943, Bibliographic and faunal index of Paleozoic pelmatozoan echinoderms: Geological Society of America Special Paper 45, 734 p.Google Scholar
Bather, F.A., 1899, A phylogenetic classification of the Pelmatozoa: British Association for the Advancement of Science, v. 1898, p. 916–923.Google Scholar
Bather, F.A., assisted by Gregory, J.W., and Goodrich, E.S., 1900, Part III, The Echinoderma, The Pelmatozoa, in Lankester, E.R., ed., A Treatise on Zoology: London, Adam and Charles Black, 344 p.Google Scholar
Bigsby, J.J., 1868, Thesaurus Siluricus; The Flora and Fauna of the Silurian Period, with Addenda (from Recent Acquisitions): London, J. Van Hoorst, lii + 214 p.Google Scholar
Billings, E., 1856, Report of progress: Canadian Naturalist and Geologist, v. 1, p. 54.Google Scholar
Billings, E., 1857a, New species of fossils from Silurian rocks of Canada: Canada Geological Survey, Report of Progress 1853−1856, Report for the Year 1856, p. 247–345.Google Scholar
Billings, E., 1857b, Fossils of the Upper Silurian rocks, Niagara and Clinton Groups: Canadian Naturalist and Geologist, ser. 1, no. 6, p. 57–60.Google Scholar
Billings, E., 1858, On the Cystidae of the Lower Silurian rocks of Canada: Canadian Organic Remains, Decade III, Canada Geological Survey, p. 9–74.Google Scholar
Billings, E., 1859, On the Crinoideae of the Lower Silurian rocks of Canada: Canadian Organic Remains, Decade 4, Canada Geological Survey, 72 p.Google Scholar
Billings, W.R., 1883, Notes on, and description of, some fossils from the Trenton Limestone: Ottawa Field Naturalists’ Club Transactions, v. 4, p. 49–52.Google Scholar
Billings, W.R., 1887, A new genus and three new species of crinoids from the Trenton Formation with notes on a large specimen of Dendrocrinus proboscidiatus : The Ottawa Naturalist (Ottawa Field Naturalists’ Club Transactions), v. 111, p. 49–54.Google Scholar
Bolton, T.E., 1970, Echinodermata from the Ordovician (Pleurocystites, Cremacrinus) and Silurian (Hemicystites, Protaxocrinus, Macnamaratylus) of Lake Timiskaming region, Ontario and Quebec, in Contributions to Canadian Paleontology: Geological Survey of Canada Bulletin, v. 187, p. 59–66.Google Scholar
Botting, J.P., 2003, Llanvirn (Middle Ordovician) echinoderms from Llandegley Rocks, central Wales: Palaeontology, v. 46, p. 685–708. doi: 10.1111/1475-4983.00316.Google Scholar
Brenchley, P.J., Carden, G.A.F., Hints, L., Kaljo, D., Marshall, J.D., Martma, T., Meidla, T., and Nolvak, J., 2003, High-resolution stable isotope stratigraphy on Upper Ordovician sequences: Constraints on the timing of bioevents and environmental changes associated with mass extinction and glaciation: Geological Society of America Bulletin, v. 115, 89–104. doi: 10.1130/0016-7606(2003)115<0089:HRSISO>2.0.CO;2.2.0.CO;2.>Google Scholar
Brett, C.E., and Brookfield, M.E., 1984, Morphology, faunas, and genesis of Ordovician hardgrounds from southern Ontario, Canada: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 46, p. 68–74.Google Scholar
Brett, C.E., and Liddell, W.D., 1978, Preservation and paleoecology of a Middle Ordovician hardground community: Paleobiology, v. 4, p. 329–348.Google Scholar
Brett, C.E., and Taylor, W.L., 1999, Middle Ordovician of the Lake Simcoe area of Ontario, Canada, in Hess, H., Ausich, W.I., Brett, C.E., and Simms, M.J. Fossil Crinoids: Cambridge, UK, Cambridge University Press, p. 68–74.Google Scholar
Brett, C.E., Deline, B.L., and McLaughlin, P.L., 2008, Attachment, facies distribution, and life history strategies in crinoids from the Upper Ordovician of Kentucky, in Ausich, W.I., and Webster, G.D., eds., Echinoderm Paleobiology: Bloomington, Indiana University Press, p. 23–52.Google Scholar
Brookfield, M.E., and Brett, C.E., 1988, Paleoenvironments of the Mid-Ordovician (upper Caradocian) Trenton limestones of southern Ontario, Canada: Storm sedimentation on a shoal-basin shelf model: Sedimentary Geology, v. 57, p. 75–105.Google Scholar
Brower, J.C., 1977, Calceocrinids from the Bromide Formation (Middle Ordovician) of southern Oklahoma: Oklahoma Geological Survey Circular 78, 28 p.Google Scholar
Brower, J.C., 1982, Phylogeny of primitive calceocrinids, in Sprinkle, J., ed., Echinoderm Faunas from the Bromide Formation (Middle Ordovician) of Oklahoma: University of Kansas Paleontological Contributions Monograph 1, p. 90–110.Google Scholar
Brower, J.C., 1988, Ontogeny and phylogeny in primitive calceocrinid crinoids: Journal of Paleontology, v. 62, p. 917–934.Google Scholar
Brower, J.C., 2007, Upper Ordovician crinoids from the Platteville Limestone of northeastern Iowa: Journal of Paleontology, v. 81, p. 103–115. doi: 10.1666/0022-3360(2007)81[103:UOCFTP]2.0.CO;2.Google Scholar
Brower, J.C., 2008, Some disparid crinoids from the Upper Ordovician (Shermanian) Walcott-Rust quarry of New York: Journal of Paleontology, v. 82, p. 57–77. doi: 10.1666/06-077.1.Google Scholar
Brower, J.C., 2011, Paleoecology of suspension-feeding echinoderm assemblages from the Upper Ordovician (Katian, Shermanian) Walcott-Rust Quarry of New York: Journal of Paleontology, v. 85, p. 369–391. doi: 10.1666/10-066.1.Google Scholar
Brower, J.C., and Strimple, H.L., 1983, Ordovician calceocrinids from northern Iowa and southern Minnesota: Journal of Paleontology, v. 57, p. 1261–1281.Google Scholar
Brower, J.C., and Veinus, J., 1974, Middle Ordovician crinoids from southwestern Virginia and eastern Tennessee: Bulletins of American Paleontology, v. 66, no. 283, 125 p.Google Scholar
Brower, J.C., and Veinus, J., 1978, Middle Ordovician crinoids from the Twin Cities area of Minnesota: Bulletins of American Paleontology, v. 74, no. 304, p. 371–506.Google Scholar
Cole, S.R., Ausich, W.I., Wright, D.F., and Koniecki, J.M., 2018, An echinoderm Lagerstätte from the Upper Ordovician (Katian), Ontario: Taxonomic re-evaluation and description of new dicyclic camerate crinoids: Journal of Paleontology, v. 92, p. 488–505. doi: 10.1017/jpa.2017.151.Google Scholar
Carpenter, P.H., 1882, On the relations of Hybocrinus, Baerocrinus, and Hybocystites : Quarterly Journal of the Geological Society of London, v. 38, p. 298–212.Google Scholar
Cuénot, L., 1948, Anatomie, éthologie et systématique des Echinodermes, Crinoides, in Grassé, P.-P., ed., Traité de Zoologie, Volume 11: Paris, Masson, p. 3–77.Google Scholar
Donovan, S.K., and Gale, A.S., 1989, Iocrinus in the Ordovician of England and Wales: Palaeontology, v. 32, p. 313–323.Google Scholar
Fritz, M.A., 1925, The stratigraphy and paleontology of Toronto and vicinity, Part 4, Hydrozoa, Echinodermata, Trilobita, and markings: Ontario Department of Mines, Annual Report for 1923, v. 32, no. 7, 34 p.Google Scholar
Grabau, A.W., and Shimer, H.W., 1910, North American Index Fossils, Invertebrates: New York, A.G. Seiler, v. 2, 909 p.Google Scholar
Grewingk, C.C.A., 1867, Über Hoplocrinus dipentas und Baerocrinus ungerni : Dorpat, Estonia, Archiv Naturkunde Live-, Ehst- und Kurlands, ser. 1, v. 4, p. 100–114.Google Scholar
Guensburg, T.E., 1984, Echinodermata of the Middle Ordovician Lebanon Limestone, central Tennessee: Bulletins of American Paleontology, v. 86, no. 319, 100 p.Google Scholar
Guensburg, T.E., 1992, Glaucocrinus falconeri Parks and Alcock, 1912 (Crinoidea) and its systematic status: Royal Ontario Museum Life Sciences Occasional Paper 39, 7 p.Google Scholar
Hall, J., 1866, Descriptions of New Species of Crinoidea and Other Fossils from the Lower Silurian Strata of the Age of the Hudson-River Group and Trenton Limestone: Albany, New York, New York State Museum of Natural History, 17 p.Google Scholar
Holland, S.M., and Patzkowsky, M.E., 1996, Sequence stratigraphy and long-term paleoceanographic change in the Middle and Upper Ordovician of the eastern United States: Geological Society of America Special Paper 306, p. 117–130.Google Scholar
Jaekel, O., 1894, Über die Morphogenie und Phylogenic der Crinoiden: Sitzungsberichten der Gesellschaft Naturforschender Freunde, Jahrgang 1894, v. 4, p. 101–121.Google Scholar
Jaekel, O., 1918, Phylogenie und System der Pelmatozoen: Paläeontologische Zeitschrift, v. 3, no. 1, 128 p.Google Scholar
Kelly, S.M., 1978, Functional morphology and evolution of Iocrinus, an Ordovician disparid inadunate crinoid [A.M. Thesis]: Bloomington, Indiana University, 80 p.Google Scholar
Kirk, E., 1914, Notes on the fossil crinoid genus Homocrinus Hall: Proceedings of the United States National Museum, v. 46, p. 473–483.Google Scholar
Kolata, D.R., 1975, Middle Ordovician echinoderms from northern Illinois and southern Wisconsin: Paleontological Society Memoir 7, Journal of Paleontology, Supplement to v. 49, no. 3, 74 p.Google Scholar
Lyon, S.S., 1869, Remarks on thirteen new species of Crinoidea from the Palaeozoic rocks of Indiana, Kentucky, and Ohio, and a description of certain peculiarities in the structure of the columns of Dolatocrinus, and their attachment to the body of the animal: Transactions of the American Philosophical Society, v. 13, p. 443–466.Google Scholar
Meek, F.B., and Worthen, A.H., 1865, Descriptions of new species of Crinoidea, etc. from the Paleozoic rocks of Illinois and some of the adjoining states: Proceedings of the Academy of Natural Sciences of Philadelphia, v. 17, p. 143–155.Google Scholar
Meek, F.B., and Worthen, A.H., 1869, Notes on some points in the structure and habits of the Palaeozoic Crinoidea: The Canadian Naturalist, n. ser., v. 4, p. 434–452.Google Scholar
Miller, J.S., 1821, A Natural History of the Crinoidea or Lily-Shaped Animals, with Observations on the Genera Asteria, Euryale, Comatula, and Marsupites : Bristol, UK, Bryan and Company, 150 p.Google Scholar
Miller, S.A., 1889, North American Geology and Paleontology: Cincinnati, Ohio, Western Methodist Book Concern, 664 p.Google Scholar
Miller, S.A., and Gurley, W.F.E., 1894, New genera and species of Echinodermata: Illinois State Museum Bulletin, v. 5, 53 p.Google Scholar
Miller, S.A., and Gurley, W.F.E., 1895, New and interesting species of Palaeozoic fossils: Illinois State Museum Bulletin, v. 7, 89 p.Google Scholar
Moore, R.C., 1962, Revision of Calceocrinidae: University of Kansas Paleontological Contributions, Echinodermata Article 4, 40 p.Google Scholar
Moore, R.C., and Lane, N.G., 1978a, Superfamily Myelodactylacea S.A. Miller, 1883, in Moore, R.C., and Teichert, K., eds., Treatise on Invertebrate Paleontology, Part T, Echinodermata 2: Boulder, Colorado, and Lawrence, Kansas, Geological Society of America (and University of Kansas Press), p. T550–T555.Google Scholar
Moore, R.C., and Lane, N.G., 1978b, Superfamily Anomalocrinacea Wachsmuth and Springer, 1886, in Moore, R.C., and Teichert, K., eds., Treatise on Invertebrate Paleontology, Part T, Echinodermata 2: Boulder, Colorado, and Lawrence, Kansas, Geological Society of America (and University of Kansas Press), p. T555–T557.Google Scholar
Moore, R.C., and Laudon, L.R., 1943, Evolution and classification of Paleozoic crinoids: Geological Society of America Special Paper 46, 151 p.Google Scholar
Moore, R.C., and Laudon, L.R., 1944, Class Crinoidea, in Shimer, H.W., and Shrock, R.R., Index Fossils of North America: New York, John Wiley and Sons, p. 137–209.Google Scholar
O’Malley, C.E., Ausich, W.I., and Chin, Y.-P., 2013, Isolation and characterization of the earliest taxon-specific organic molecules (Mississippian, Crinoidea): Geology, v. 41, p. 347–350. doi: 10.1130/G33792.1.Google Scholar
Parks, W.A., 1908, On an occurrence of Hybocystis in Ontario: Ottawa Naturalist, v. 21, p. 232–236.Google Scholar
Parks, W.A., and Alcock, F.J., 1912, On two new crinoids from the Trenton Formation of Ontario: Ottawa Naturalist, v. 26, p. 41–45.Google Scholar
Parsley, R.L., 1981, Echinoderms from Middle and Upper Ordovician rocks of Kentucky: United States Geological Survey Professional Paper, 1066-K, p. 1–9.Google Scholar
Peters, S.E., and Ausich, W.I., 2008, A sampling-adjusted macroevolutionary history for Ordovician-Early Silurian crinoids: Paleobiology, v. 34, p. 104–116. doi: 10.1666/07035.1.Google Scholar
Ramsbottom, W.H.C., 1961, A monograph on British Ordovician Crinoidea: Palaeontographical Society, v. 114, p. 1–37.Google Scholar
Regnell, G., 1948, Swedish Hybocrinida (Crinoidea Inadunata Disparata; Ordovician−Lower Silurian): Arkiv for Zoologi, v. 40, no. 9, p. 1–27.Google Scholar
Ringueberg, E.N.S., 1889, The Calceocrinidae. A revision of the family, with descriptions of some new species: Annals of the New York Academy of Science, v. 4, p. 388–408.Google Scholar
Rozhnov, S.V., 1985, Morfologiya, simmetriya i sistematicheskoe polozhenie morskikh lilii gibokrinid [Morphology, symmetry and systematic position of the hybocrinid crinoids]: Paleontologicheskii Zhurnal, no. 2, p. 4–16.Google Scholar
Rozhnov, S.V., 1986, Morphology, symmetry, and systematic position of the Hybocrinida (Crinoidea): Paleontological Journal, v. 19, no. 2, p. 1–13.Google Scholar
Rozhnov, S.V., 2002, Morphogenesis and evolution of crinoids and other pelmatozoan echinoderms in the early Paleozoic: Paleontological Journal, v. 36, Supplement 6, p. S525–S674.Google Scholar
Sardeson, F.W., 1928, Derivation of the Calceocrinidae: Pan-American Geologist, v. 49, p. 35–46.Google Scholar
Sheehan, P.M., 2001, The Late Ordovician mass extinction: Annual Reviews of Earth and Planetary Sciences, v. 29, p. 331–364. doi: 10.1146/annurev.earth.29.1.331.Google Scholar
Shumard, B.F., 1868 (‘1866’), A catalogue of the Palaeozoic fossils of North America, Part I, Paleozoic Echinodermata: Transactions of the Saint Louis Academy of Science, v. 2, p. 334–407.Google Scholar
Sinclair, G. W., 1945, The age of the Ordovician Kirkfield Formation in Ontario: Ohio Journal of Science, v. 54, p. 31–41.Google Scholar
Sloan, R.E., 2005, Minnesota Fossils and Fossiliferous Rocks: Winona, Minnesota, privately published, 218 p.Google Scholar
Smith, A.B., 1985, Crinoidea, in Murray, J.W., ed., Atlas of Invertebrate Macrofossils: Harlow, Essex, UK, The Palaeontological Association, Longman, p. 153–181.Google Scholar
Springer, F., 1911, On a Trenton echinoderm fauna: Canada Department Mines, Memoir 15-P, 70 p.Google Scholar
Springer, F., 1926, American Silurian Crinoids: Smithsonian Institution Publication, v. 2872, 239 p.Google Scholar
Sprinkle, J., 1973, Morphology and evolution of blastozoan echinoderms: Cambridge, Massachusetts, Harvard University Museum of Comparative Zoology, Special Publication, 283 p.Google Scholar
Sprinkle, J., and Moore, R.C., 1978, Hybocrinida, in Moore, R.C., and Teichert, K., eds., Treatise on Invertebrate Paleontology, Part T, Echinodermata 2: Boulder, Colorado, and Lawrence, Kansas, Geological Society of America (and University of Kansas Press), p. T564–T575.Google Scholar
Sproat, C.D., Jin, J., Zhan, R.B., and Rudkin, D.M., 2015, Morphological variability and paleoecology of the Late Ordovician Parastrophina from eastern Canada and the Tarim Basin, Northwest China: Palaeoworld, v. 24, p. 160–175. doi: 10.1016/j.palwor.2014.11.008.Google Scholar
Steele, H.M., and Sinclair, G.W., 1971, A Middle Ordovician fauna from Braeside, Ottawa Valley, Ontario: Geological Survey Canada Bulletin, v. 211, p. 97 p.Google Scholar
Strimple, H.L., 1975, Bottom-dwelling hybocrinids from Kentucky: Southeastern Geology, v. 17, no. 1, p. 51–53.Google Scholar
Strimple, H.L., and Watkins, W.T., 1949, Hybocrinus crinerensis, new species from the Ordovician of Oklahoma: American Journal of Science, v. 247, p. 131–133.Google Scholar
Ubaghs, G., 1953, Classe des Crinoides, in Piveteau, J., ed., Traité de Paléontologie. Volume 3: Paris, Masson & Cie, p. 658–773.Google Scholar
Ubaghs, G., 1978, Skeletal morphology of fossil crinoids, in Moore, R.C., and Teichert, K., eds., Treatise on Invertebrate Paleontology, Part T, Echinodermata 2: Boulder, Colorado, and Lawrence, Kansas, Geological Society of America (and University of Kansas Press), p. T58–T216.Google Scholar
Ulrich, E.O., 1886, Remarks upon the names Cheirocrinus and Calceocrinus, with descriptions of three new generic terms and one new species: Minnesota Geology and Natural History Survey, Annual Report 14, p. 104–113.Google Scholar
Ulrich, E.O., 1925, New classification of the ‘Heterocrinidae, in Foerste, A.F. Upper Ordovician Faunas of Ontario and Quebec: Canada Geological Survey Memoir 138, p. 82–106.Google Scholar
Wachsmuth, C., and Springer, F., 1880–1886, Revision of the Palaeocrinoidea: Proceedings of the Academy of Natural Sciences of Philadelphia, Part 1, The families Ichthyocrinidae and Cyathocrinidae (1880), p. 226–378 (separate repaged p. 1–153); Part 2, Family Sphaeroidocrinidae, with the sub-families Platycrinidae, Rhodocrinidae, and Actinocrinidae (1881), p. 177–411 (separate repaged p. 1–237); Part 3, Sec. 1. Discussion of the classification and relations of the brachiate crinoids, and conclusion of the generic descriptions (1885), p. 225–364 (separate repaged 1–138); Part 3, Sec. 2, Discussion of the classification and relations of the brachiate crinoids, and conclusion of the generic descriptions (1886), p. 64–226 (separately repaged p. 139–302).Google Scholar
Wachsmuth, C., and Springer, F., 1891, The perisomic plates of the crinoids: Proceedings of the Academy of Natural Sciences of Philadelphia, v. 41, p. 345–392.Google Scholar
Walcott, C.D., 1883, Descriptions of new species of fossils from the Trenton Group of New York: New York State Museum of Natural History, Annual Report, v. 35, p. 207–214 (advance publication).Google Scholar
Walcott, C.D., 1884, Descriptions of new species of fossils from the Trenton Group of New York: New York State Museum of Natural History, Annual Report, v. 35, p. 207–214.Google Scholar
Warn, J.M., and Strimple, H.L., 1977, The disparid inadunate superfamilies Homocrinacea and Cincinnaticrinacea (Echinodermata. Crinoidea), Ordovician-Silurian, North America: Bulletins of American Paleontology, v. 72, no. 296, p. 1–13.Google Scholar
Webby, G.D., Paris, F., Droser, M.L., and Percival, I.G., 2004, The Great Ordovician Biodiversification Event: New York, Columbia University Press, 484 p.Google Scholar
Webster, G.D., 1973, Bibliography and index of Paleozoic crinoids, 1942–1968: Geological Society of America Memoir 137, 341 p.Google Scholar
Webster, G.D., 1986, Bibliography and index of Paleozoic crinoids, 1974–1980: Geological Society of America Microform Publication 16, 405 p., 5 cards.Google Scholar
Webster, G.D., 1988, Bibliography and index of Paleozoic crinoids and coronate echinoderms 1981–1985: Geological Society of America Microform Publication, v. 18, 235 p., 3 cards.Google Scholar
Webster, G.D., 1993, Bibliography and index of Paleozoic crinoids, 1986–1990: Geological Society of America Microform Publication 25, p. 204 p., 3 cards.Google Scholar
Webster, G.D., and Webster, D.W., 2013, Bibliography and Index of Paleozoic Crinoids, Coronates, and Hemistreptocrinoids, 1758–2012: http://crinoids.azurewebsites.net/ (accessed 04 April 2016).Google Scholar
Wetherby, A.G., 1880, Remarks on the Trenton Limestone of Kentucky, with descriptions of new fossils from that formation and the Kaskaskia (Chester) Group, sub-Carboniferous: Journal of the Cincinnati Society of Natural History, v. 3, p. 144–160.Google Scholar
Wilson, A.E., 1946, Echinodermata of the Ottawa Formation of the Ottawa-St. Lawrence lowland: Canada Geological Survey Bulletin, v. 4, p. 1–61.Google Scholar
Wright, D.F., 2017, Bayesian estimation of fossil phylogenies and the evolution of early to middle Paleozoic crinoids (Echinodermata): Journal of Paleontology, v. 91, p. 799–814. doi: 10.1017/jpa.2016.141.Google Scholar
Wright, D.F., and Toom, U., 2017, New crinoids from the Baltic region (Estonia): Fossil tip-dating phylogenetics constrains the origin and Ordovician-Silurian diversification of the Flexibilia (Echinodermata): Palaeontology, v. 60, p. 893–910. doi: 10.1111/pala.12324.Google Scholar
Wright, D.F., Ausich, W.I., Cole, S.R., Rhenberg, E.C., and Peter, M.E., 2017, Phylogenetic taxonomy and classification of the Crinoidea (Echinodermata): Journal of Paleontology, v. 91 p. 829–846. doi: 10.1017/jpa.2016.142.Google Scholar
Yakovlev, N.N., 1964, Klass Crinoidea, Mosskie lilii, Obstchaya chast, Osnovy paleontologii [Class Crinoidea, crinoids, general part, characteristics, and morphology], in Orlov, Yu.A., ed., Spravotchnik dlya Paleontologov i Geologov SSR, Echinodermata, Hemichordata, Pogonophora, and Chaetognatha [Fundamentals of Paleontology, Echinodermata, Hemichordata, Pogonophora, and Chaetognatha]: Moscow, Izdatel’stvo Nedra, p. 54–74.Google Scholar
Zamora, S., Rahman, I.A., and Ausich, W.I., 2015, A new iocrinid crinoid (Disparida) from the Ordovician (Darriwilian) of Morocco: PeerJ, v. 3, p. e1450, 19 p. doi: 10.7717/peerj.14500.Google Scholar
Zittel, K.A. von, 1879, Handbuch der Palaeontologie, Volume 1, Palaeozoologie, No. 1: Munich, Germany, R. Oldenbourg, 765 p. (Echinoderms, p. 308–560).Google Scholar