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Faunal overview of the Mud Hill locality from the early Permian Vale Formation of Taylor County, Texas

Published online by Cambridge University Press:  11 July 2018

Bryan M. Gee
Affiliation:
Department of Biology, University of Toronto Mississauga, Ontario L5L 1C6, Canada 〈bryan.gee@mail.utoronto.ca〉 〈diane.scott@utoronto.ca〉 〈robert.reisz@utoronto.ca〉
Steven J. Rosscoe
Affiliation:
Department of Geology, Hardin-Simmons University, Abilene, Texas 79601, USA 〈srosscoe@hsutx.edu〉
Diane Scott
Affiliation:
Department of Biology, University of Toronto Mississauga, Ontario L5L 1C6, Canada 〈bryan.gee@mail.utoronto.ca〉 〈diane.scott@utoronto.ca〉 〈robert.reisz@utoronto.ca〉
Judie Ostlien
Affiliation:
Fossil Finder, Olhausen Estate, Abilene, Texas 79606, USA 〈brianbethel72@gmail.com〉
Robert R. Reisz
Affiliation:
Department of Biology, University of Toronto Mississauga, Ontario L5L 1C6, Canada 〈bryan.gee@mail.utoronto.ca〉 〈diane.scott@utoronto.ca〉 〈robert.reisz@utoronto.ca〉

Abstract

The Texas red beds represent one of the richest series of early Permian deposits in the world. In particular, the Clear Fork Group has produced a diverse assemblage of temnospondyls, early reptiles, and synapsids. However, most of this material has been sourced from the oldest member, the Arroyo Formation, and the understanding of the paleoecosystem of the younger Vale and Choza formations is less well resolved. Here we present a previously undescribed Vale locality, the first vertebrate-bearing locality from the formation to be described in detail in several decades, from near Abilene, Texas with juvenile diplocaulids, captorhinids, abundant material of rare taxa such as Varanops and diadectids, and the first report of a recumbirostran ‘microsaur’ from the formation. This assemblage is atypical of early Permian deposits in the taxonomic and size distribution of the vertebrate fauna in comparison to other localities from the Vale Formation that preserve a greater abundance of aquatic taxa (e.g., fishes, Trimerorhachis) and synapsids (e.g., Dimetrodon). Minimal abrasion of the elements, relative articulation and association of the specimen of Varanops, and the paucity of aquatic taxa suggest an ephemeral pond deposit in which organisms were preserved essentially in situ. Our characterization of the locality also permits a revision and discussion of the vertebrate faunal assemblage of the Vale Formation.

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Articles
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Copyright © 2018, The Paleontological Society 

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References

Anderson, J.S., 2007, Incorporating ontogeny into the matrix: A phylogenetic evaluation of developmental evidence for the origins of modern amphibians, in Anderson, J.S., and Sues, H.-D., eds., Major Transitions in Vertebrate Evolution: Bloomington, Indiana University Press, p. 182227.Google Scholar
Anderson, J.S., Scott, D., and Reisz, R.R., 2009, Nannaroter mckinziei, a new ostodolepid ‘microsaur’ (Tetrapoda, Lepospondyli, Recumbirostra) from the early Permian of Richards Spur (Ft. Sill), Oklahoma: Journal of Vertebrate Paleontology, v. 29, p. 379388. doi: 10.1671/039.029.0222.Google Scholar
Beerbower, J.R., 1963, Morphology, paleo-ecology and phylogeny of the Permo-Pennsylvanian amphibian Diploceraspis : Bulletin of the Museum of Comparative Zoology, v. 130, p. 31108.Google Scholar
Behrensmeyer, A.K., 1978, Taphonomic and ecologic information from bone weathering: Paleobiology, v. 4, p. 150162.Google Scholar
Berman, D.S., Henrici, A.C., Kissel, R.A., Sumida, S.S., and Martens, T., 2004, A new diadectid (Diadectomorpha), Orobates pabsti, from the early Permian of central Germany: Bulletin of Carnegie Museum of Natural History, v. 35, p. 136. doi: 10.2992/0145-9058(2004)35[1:ANDDOP]2.0.C);2.Google Scholar
Beyrich, E., 1848, Über Xenacanthus decheni und Holacanthus gracilis, zwei Fische aus der Formation des Rothliegenden in Nord Deutschland: Berlin, Preussische Akademie der Wissenschaften, v. 1848, p. 2433.Google Scholar
Bolt, J.R., and Rieppel, O., 2009, The holotype skull of Llistrofus pricei Carroll and Gaskill, 1978 (Microsauria: Hapsidopareiontidae): Journal of Paleontology, v. 83, p. 471483. doi: 10.1666/08-076.1.Google Scholar
Bossy, K.A., and Milner, A.C., 1998, Order Nectridea, in Wellnhofer, P., ed., Encyclopedia of Paleoherpetology, Part 1, Lepospondyli, Munich, Verlag Dr. Friedrich Pfeil, p. 73132.Google Scholar
Broili, F., 1904, Permische Stegocephalen und Reptilien aus Texas: Zweite Hälfte: Palaeontolographica, v. 1904, p. 49120.Google Scholar
Campione, N.E., and Reisz, R.R., 2010, Varanops brevirostris (Eupelycosauria: Varanopidae) from the lower Permian of Texas, with discussion of varanopid morphology and interrelationships: Journal of Vertebrate Paleontology, v. 30, p. 724746. doi: 10.1080/02724631003762914.Google Scholar
Carroll, R.L., and Gaskill, P., 1978, The order Microsauria: Memoirs of the American Philosophical Society, v. 126, p. 1211.Google Scholar
Case, E.C., 1907, Revision of the Pelycosauria of North America: Carnegie Institution of Washington Publication, v. 55, 176 p.Google Scholar
Case, E.C., 1910, New or little known reptiles and amphibians from the Permian of Texas: Bulletin of the American Museum of Natural History, v. 28, p. 163181.Google Scholar
Case, E.C., 1911, A revision of the Cotylosauria of North America: Carnegie Institution of Washington Publication, v. 145, p. 1122.Google Scholar
Chaney, D.S., Sues, H.-D., and DiMichele, W.A., 2005, A juvenile skeleton of the nectridean amphibian Diplocaulus and associated flora and fauna from the Mitchell Creek Flats locality (upper Waggoner Ranch Formation; early Permian), Baylor County, north-central Texas, USA: New Mexico Museum of Natural History and Science Bulletin, v. 30, p. 3947.Google Scholar
Cope, E.D., 1868, Synopsis of the extinct Batrachia of North America: Proceedings of the Academy of Natural Sciences of Philadelphia, v. 20, p. 208221.Google Scholar
Cope, E.D., 1877a, A continuation of researches among the Batrachia of the coal measures of Ohio: Proceedings of the American Philosophical Society, v. 16, p. 573578.Google Scholar
Cope, E.D., 1877b, Descriptions of extinct Vertebrata from the Permian and Triassic formations of the United States: Proceedings of the American Philosophical Society, v. 17, p. 182193.Google Scholar
Cope, E.D., 1878, Descriptions of extinct Batrachia and Reptilia from the Permian formation of Texas: Proceedings of the American Philosophical Society, v. 17, p. 505530.Google Scholar
Cope, E.D., 1880, Second contribution to the history of the Vertebrata of the Permian formation of Texas: Proceedings of the American Philosophical Society, v. 19, p. 3858.Google Scholar
Cope, E.D., 1881, Catalogue of Vertebrata of the Permian formation of the United States: American Naturalist, v. 15, p. 162164.Google Scholar
Cope, E.D., 1882, Third contribution to the history of the Vertebrata of the Permian formation of Texas: Proceedings of the American Philosophical Society, v. 20, p. 447461.Google Scholar
Cope, E.D., 1883, Fourth contribution to the history of the Permian formation of Texas: Paleontological Bulletin, v. 36, p. 628636.Google Scholar
Cope, E.D., 1884, On the structure of the skull in the elasmobranch Didymodus : Proceedings of the American Philosophical Society, v. 21, p. 572590.Google Scholar
Cope, E.D., 1895, The reptilian order Cotylosauria: Proceedings of the American Philosophical Society, v. 34, p. 436457.Google Scholar
Cope, E.D., 1896, Second contribution to the history of the Cotylosauria: Proceedings of the American Philosophical Society, v. 34, p. 436457.Google Scholar
Dalquest, W.W., and Mamay, S.H., 1963, A remarkable concentration of Permian amphibian remains in Haskell County, Texas: Journal of Geology, v. 71, p. 641644.Google Scholar
Daly, E., 1973, A lower Permian vertebrate fauna from southern Oklahoma: Journal of Paleontology, v. 47, p. 562589.Google Scholar
Dawson, J.W., 1863, Air-breathers of the Coal Period: A Descriptive Account of the Remains of Land Animals Found in the Coal Formation of Nova Scotia: Montréal, Dawson Brothers, 118 p.Google Scholar
de Ricqlès, A., and Taquet, P., 1982, La faune de vertebres de Permien supérieur du Niger, I, Le captorhinomorphe Moradisaurus grandis (Reptilia, Cotylosauria)—le crane: Annales de Paléontologie, v. 68, p. 3363.Google Scholar
Flear, V.J., Modesto, S.P., and Reisz, R.R., 2017, New diadectomorph cotylosaur material from the middle Clear Fork Formation, lower Permian of Texas: Journal of Vertebrate Paleontology, v. 37, supplement 1, p. 113.Google Scholar
Fox, R.C., and Bowman, M.C., 1966, Osteology and relationships of Captorhinus aguti (Cope) (Reptilia: Captorhinomorpha): University of Kansas Paleontological Contributions, v. 15, p. 180.Google Scholar
Fritsch, A., 1876, Über die Fauna der Gaskohle des Pilsner und Rakonitzer Beckens: Sitzungsberichte der Böhmischen Gesellschaft der Wissenschaften Prague, v. 1875, p. 7079.Google Scholar
Germain, D., 2010, The Moroccan diplocaulid: The last lepospondyl, the single one on Gondwana: Historical Biology, v. 22, p. 439. doi: 10.1080/08912961003779678.Google Scholar
Goodrich, E.S., 1930, Studies on the Structure and Development of Vertebrates: New York, Dover Publications, 837 p.Google Scholar
Haglund, T.R., 1977, New occurrences and paleoecology of Peronedon primus Olson (Nectridea): Journal of Paleontology, v. 51, p. 982985.Google Scholar
Hentz, T.F., 1988, Lithostratigraphy and paleoenvironments of upper Paleozoic continental red beds, north-central Texas—Bowie (new) and Wichita (revised) groups: Bureau of Economic Geology, University of Texas at Austin, Report of Investigations, v. 170, 55 p.Google Scholar
Hunt, A.P., and Lucas, S.G., 2005, The origin of large vertebrate coprolites from the early Permian of Texas: New Mexico Museum of Natural History and Science Bulletin, v. 30, p. 125126.Google Scholar
Jaekel, O., 1903, Über Ceraterpeton, Diceratosaurus, und Diplocaulus : Neues Jahrbuch für Geologie und Palaontologie, v. 1903, p. 109134.Google Scholar
Kemp, T.S., 1982, Mammal-like Reptiles and the Origin of Mammals: London, Academic Press, 363 p.Google Scholar
LeBlanc, A.R., and Reisz, R.R., 2013, Periodontal ligament, cementum, and alveolar bone in the oldest herbivorous tetrapods, and their evolutionary significance: PLOS One, v. 8, p. e74697 doi: 10.1371/journal.pone.0074697.Google Scholar
Lucas, S.G., 2006, Global Permian tetrapod biostratigraphy and biochronology: Geological Society: London, Special Publication, v. 265, p. 6593. doi: 10.1144.GSL.SP.2006.265.01.04.Google Scholar
Lucas, S.G., and Hunt, A.P., 2005, Permian tetrapod tracks from Texas: New Mexico Museum of Natural History and Science Bulletin, v. 30, p. 202206.Google Scholar
Maddin, H.C., Evans, D.C., and Reisz, R.R., 2006, An early Permian varanodontine varanopid (Synapsida: Eupelycosauria) from the Richards Spur locality, Oklahoma: Journal of Vertebrate Paleontology, v. 26, p. 957966. doi: 10.1671/0272-4634(2006)26[957:AEPVVS]2.0.C0;2.Google Scholar
Marsh, O.C., 1878, Notice of new fossil reptiles: American Journal of Science, v. 89, p. 409411.Google Scholar
May, W.J., and Hall, J.D., 2016, First occurrence of the diplocaulid genus Diploceraspis from the Wellington Formation (lower Permian) of northern Oklahoma: Transactions of the Kansas Academy of Science, v. 119, p. 193199. doi: 10.1660/062.119.0209.Google Scholar
Mead, J.G., 1971, Paleontology of the Sid Mc-Adams locality (Permian) southern Taylor County, Texas [MS thesis]: Austin, University of Texas, 62 p.Google Scholar
Miall, L.C., 1875, Report of the committee on the structure and classification of the labyrinthodonts: Report of the Meeting of the British Association for the Advancement of Science, v. 1874, p. 149192.Google Scholar
Milner, A.C., 1985, On the identity of Trematopsis seltini (Amphibia: Temnospondyli) from the lower Permian of Texas: Neues Jahrbuch für Geologie und Palaontologie, Monatshefte, v. 1985, p. 357367.Google Scholar
Milner, A.C., and Ruta, M., 2009, A revision of Scincosaurus (Tetrapoda, Nectridea) from the Moscovian of Nyrany, Czech Republic, and the phylogeny and interrelationships of nectrideans: Special Papers in Palaeontology, v. 81, p. 7189. doi: 10.1111/j.1475-4983.2009.00883.x.Google Scholar
Modesto, S.P., Scott, D.M., Berman, D.S., Müller, J., and Reisz, R.R., 2007, The skull and the palaeoecological significance of Labidosaurus hamatus, a captorhinid reptile from the lower Permian of Texas: Zoological Journal of the Linnean Society, v. 149, p. 237262. doi: 10.1111/j.1096-3642.2007.00242.x.Google Scholar
Modesto, S.P., Lamb, A.J., and Reisz, R.R., 2014, The captorhinid reptile Captorhinikos valensis from the lower Permian Vale Formation of Texas, and the evolution of herbivory in eureptiles: Journal of Vertebrate Paleontology, v. 34, p. 291302. doi: 10.1080/02724634.2013.809358.Google Scholar
Modesto, S.P., Flear, V.J., Dilney, M.M., and Reisz, R.R., 2016, A large moradisaurine tooth plate from the lower Permian of Texas and its biostratigraphic implications: Journal of Vertebrate Paleontology, v. 36, p. e1221832 doi: 10.1080/02724634.2016.1221832.Google Scholar
Nelson, J.W., Hook, R., and Chaney, D., 2013, Lithostratigraphy of the lower Permian (Leonardian) Clear Fork Formation of north-central Texas: New Mexico Museum of Natural History and Science Bulletin, v. 60, p. 286311.Google Scholar
Olson, E.C., 1939, The fauna of the Lysorophus pockets in the Clear Fork Permian, Baylor County, Texas: Journal of Geology, v. 47, p. 389397.Google Scholar
Olson, E.C., 1947, The family Diadectidae and its bearing on the classification of reptiles: Fieldiana Geology, v. 11, p. 153.Google Scholar
Olson, E.C., 1948, A preliminary report on vertebrates from the Permian Vale Formation of Texas: The Journal of Geology, v. 56, p. 186198.Google Scholar
Olson, E.C., 1951a, Diplocaulus: a study in growth and variation: Fieldiana Geology, v. 11, p. 57154.Google Scholar
Olson, E.C., 1951b, Fauna of Upper Vale and Choza: 1−5: Fieldiana Geology, v. 10, p. 89128.Google Scholar
Olson, E.C., 1951c, Vertebrates from the Choza Formation, Permian of Texas: The Journal of Geology, v. 59, p. 178181.Google Scholar
Olson, E.C., 1952a, Fauna of the Upper Vale and Choza: 6, Diplocaulus : Fieldiana Geology, v. 10, p. 147166.Google Scholar
Olson, E.C., 1952b, The evolution of a Permian vertebrate chronofauna: Evolution, v. 6, p. 181196.Google Scholar
Olson, E.C., 1954a, Fauna of the Vale and Choza: 7, Pelycosauria, family Caseidae: Fieldiana Geology, v. 10, p. 193204.Google Scholar
Olson, E.C., 1954b, Fauna of the Vale and Choza: 8, Pelycosauria, Dimetrodon : Fieldiana Geology, v. 10, p. 205210.Google Scholar
Olson, E.C., 1954c, Fauna of the Vale and Choza: 9, Captorhinomorpha: Fieldiana Geology, v. 10, p. 211218.Google Scholar
Olson, E.C., 1955, Fauna of the Vale and Choza: 10, Trimerorhachis, including a revision of pre-Vale species: Fieldiana Geology, v. 10, p. 225274.Google Scholar
Olson, E.C., 1956a, Fauna of the Vale and Choza: 11, Lysorophus, Vale and Choza, Diplocaulus, Cacops, and Eryopidae, Choza: Fieldiana Geology, v. 10, p. 312322.Google Scholar
Olson, E.C., 1956b, Fauna of the Vale and Choza: 12, a new trematopsid amphibian from the Vale Formation: Geology, v. 10, p. 323328.Google Scholar
Olson, E.C., 1956c, Fauna of the Vale and Choza: 13, Diadectes, Xenacanthus, and specimens of uncertain affinities: Fieldiana Geology, v. 10, p. 329334.Google Scholar
Olson, E.C., 1958, Fauna of the Vale and Choza: 14, summary, review, and integration of the geology and the faunas: Fieldiana Geology, v. 10, p. 397448.Google Scholar
Olson, E.C., 1965, New Permian vertebrates from the Chickasha Formation in Oklahoma: Oklahoma Geological Survey Circular, v. 70, p. 170.Google Scholar
Olson, E.C., 1970, New and little known genera and species of vertebrates from the lower Permian of Oklahoma: Fieldiana Geology, v. 18, p. 359434.Google Scholar
Olson, E.C., 1974, On the source of therapsids: Annals of the South African Museum, v. 64, p. 2746.Google Scholar
Olson, E.C., 1979a, Aspects of the biology of Trimerorhachis (Amphibia: Temnospondyli): Journal of Paleontology, v. 53, p. 117.Google Scholar
Olson, E.C., 1979b, Seymouria grandis n. sp. (Batrachosauria: Amphibia) from the middle Clear Fork (Permian) of Oklahoma and Texas: Journal of Paleontology, v. 53, p. 720728.Google Scholar
Olson, E.C., 1989, The Arroyo Formation (Leonardian: lower Permian) and its vertebrate fossils: Texas Memorial Museum Bulletin, v. 35, p. 125.Google Scholar
Olson, E.C., and Mead, J.G., 1982, The Vale Formation (lower Permian): Its vertebrates and paleoecology: Texas Memorial Museum Bulletin, v. 29, p. 146.Google Scholar
Osborn, H.F., 1903, The reptilian subclasses Diapsida and Synapsida and the early history of the Diaptosaura: Memoirs of the American Museum of Natural History, v. 1, p. 449508.Google Scholar
Pardo, J.D., and Anderson, J.S., 2016, Cranial morphology of the Carboniferous-Permian tetrapod Brachydectes newberryi (Lepospondyli, Lysorophia): New data from µCT: PLOS One, v. 11, p. e0161823. doi: 10.1371/journal.pone.0161823.Google Scholar
Reisz, R.R., and Tsuji, L.A., 2006, An articulated skeleton of Varanops with bite marks: The oldest known evidence of scavenging among terrestrial vertebrates: Journal of Vertebrate Paleontology, v. 26, p. 10211023. doi: 10.1671/0272-4634(2006)26[1021:AASOVW]2.0.CO;2.Google Scholar
Reisz, R.R., Liu, J., Li, J.L., and Müller, J., 2011, A new captorhinid reptile, Gansurhinus qingtoushanensis, gen. et sp. nov., from the Permian of China: Naturwissenschaften, v. 98, p. 435441. doi: 10.1007/s00114-011-0793-0.Google Scholar
Romer, A.S., and Price, L.I., 1940, Review of the Pelycosauria: Geological Society of America: Special Paper, v. 28, p. 1538.Google Scholar
Schultze, H.-P., and Foreman, J.R., 1981, A new gymnarthrid microsaur from the lower Permian of Kansas with a review of the tuditanomorph microsaurs (Amphibia): Occasional Papers of the Museum of Natural History, University of Kansas, v. 91, p. 125.Google Scholar
Shumard, B.F., 1858, Notice of new fossils from the Permian strata of New Mexico and Texas, collected by Dr. George G. Shumard, geologist for the United States government expedition for obtaining water by means of artesian wells along the 32nd parallel, under the direction of Captain John Pope, US Topographic Engineers: St. Louis Academy of Science Transactions, v. 1, p. 290297.Google Scholar
Shumard, B.F., 1859, Fossils from the Permian strata of Texas and New Mexico: Transactions of the St. Louis Academy of Science, v. 1, p. 387403.Google Scholar
Stovall, J.W., 1950, A new cotylosaur from north central Oklahoma: American Journal of Science, v. 248, p. 4654.Google Scholar
Sumida, S.S., 1989, The appendicular skeleton of the early Permian genus Labidosaurus (Reptilia, Captorhinomorpha, Captorhinidae) and the hind limb musculature of captorhinid reptiles: Journal of Vertebrate Paleontology, v. 9, p. 295313.Google Scholar
Szostakiwskyj, M., Pardo, J.D., and Anderson, J.S., 2015, Micro-CT study of Rhynchonkos stovalli (Lepospondyli, Recumbirostra), with description of two new genera: PLOS One, v. 10, p. e0127307 doi: 10.1371/journal.pone.0127307.Google Scholar
Tabor, N.J., and Montañez, I.P., 2002, Shifts in late Paleozoic atmospheric circulation over western equatorial Pangea: Insights from pedogenic mineral δ18O compositions: Geology, v. 30, p. 11271130. doi: 10.1130/0091-7613(2002)030<1127:SILPAC>2.0.CO;2.2.0.CO;2.>Google Scholar
Tabor, N.J., Montañez, I.P., and Southard, R.J., 2002, Paleoenvironmental reconstruction from chemical and isotopic compositions of Permo-Pennsylvanian pedogenic minerals: Geochimica et Cosmochimica Acta, v. 66, p. 30933107. doi: 10.1016/S0016-7037(02)00879-7.Google Scholar
Tabor, N.J., Montañez, I.P., Scotese, C.R., Poulsen, C.J., Mack, G.H., Fielding, C.R., Frank, T.D., and Isbell, J.L., 2008, Paleosol archives of environmental and climatic history in paleotropical western Pangea during the latest Pennsylvanian through early Permian: Resolving the late Paleozoic ice age in time and space: Geological Society of America Special Paper, v. 441, p. 291303. doi: 10.1130/2008.2441(20).Google Scholar
Vaugh, P.P., 1958, A specimen of the captorhinid reptile Captorhinikos chozaensis Olson, 1954, from the Hennessey Formation, lower Permian of Oklahoma: Journal of Geology, v. 66, p. 327332.Google Scholar
Watson, D.M.S., 1917, A sketch classification of the pre-Jurassic tetrapod vertebrates: Proceedings of the Zoological Society of London, v. 1917, p. 167186.Google Scholar
Williston, S.W., 1909a, New or little-known Permian vertebrates: Trematops, new genus: Journal of Geology, v. 17, p. 636658.Google Scholar
Williston, S.W., 1909b, The skull and extremities of Diplocaulus : Transactions of the Kansas Academy of Science, v. 22, p. 122131.Google Scholar
Williston, S.W., 1910a, Cacops, Desmospondylus: New genera of Permian vertebrates: Geological Society of America Bulletin, v. 21, p. 249284.Google Scholar
Williston, S.W., 1910b, New Permian reptiles: Rhachitomous vertebrae: Journal of Geology, v. 18, p. 585600.Google Scholar
Williston, S.W., 1911, American Permian Vertebrates: Chicago, University of Chicago Press, 145 p.Google Scholar
Williston, S.W., 1913, Ostodolepis brevispinatus, a new reptile from the Permian of Texas: Journal of Geology, v. 21, p. 363366.Google Scholar
Williston, S.W., 1914, The osteology of some American Permian vertebrates: Journal of Geology, v. 22, p. 364419.Google Scholar
Wilson, J.A., 1948, Vertebrate fauna of a Permian locality in Taylor County, Texas: Bulletin of the Geological Society of America (Abstracts), v. 29, p. 1363.Google Scholar
Wilson, J.A., 1953, Permian vertebrates from Taylor County, Texas: Journal of Paleontology, v. 27, p. 456470.Google Scholar
Woodhead, J., Reisz, R., Fox, D., Drysdale, R., Hellstrom, J., Maas, R., Cheng, H., and Edwards, R.L., 2010, Speleothem climate records from deep time? Exploring the potential with an example from the Permian: Geology, v. 38, p. 455458. doi: 10.1130/G30354.1.Google Scholar
Zittel, K.V., 1888, Handbuch der Palaeontogie, 1, Abt. Palaeozoologie, III Band, Amphibia: Munich, Oldenbourg, 971 p.Google Scholar