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A quantitative review of European stenonoid horses

Published online by Cambridge University Press:  20 May 2016

M. T. Alberdi
Affiliation:
Departamento de Paleobiología, Museo Nacional de Ciencias Naturales, CSIC, José Gutiérrez Abascal, 2, 28006 Madrid, Spain
E. Ortiz-Jaureguizar
Affiliation:
Departamento de Paleobiología, Museo Nacional de Ciencias Naturales, CSIC, José Gutiérrez Abascal, 2, 28006 Madrid, Spain Laboratorio de Sistemática y Biología Evolutiva (LASBE), Museo de La Plata, Paseo del Bosque s/n, 1900 La Plata, Argentina
J. L. Prado
Affiliation:
INCUAPA, Facultad Ciencias Sociales, UNC, Provincia de Buenos Aires, San Martin 3060, 7400 Olavarría, Argentina

Abstract

The taxonomy of the European stenonoid horses is revised using multivariate statistical techniques in this paper. The objectives are: to identify the taxonomic status of this group; to revise the specific and subspecific diagnoses, and the valid taxa; and to analyze their paleoecology and evolution. We conclude that: 1) there are six species among the Plio-Pleistocene European stenonoids: Equus major Depéret, in Delafond and Depéret 1893, ex Boule; Equus livenzovensis Bajgusheva, 1978; Equus stenonis Cocchi, 1867; Equus senezensis Prat, 1964; Equus altidens von Reichenau, 1915; and Equus sussenbornensis Wüts, 1901; 2) There are seven subspecies, three of E. stenonis, two of E. senezensis, and two of E. altidens; 3) There are two monophyletic groups according to the “neighbor-joining method,” one composed of the middle-to-small-size species, and the other of the biggest ones; 4) There is a trend toward decreasing body size through time in stenonoid horses. This trend, that follows the inverse direction of Cope's rule, can be related to the climatic-environmental changes that occurred from the middle Villafranchian to the early Galerian; 5) Large stenonoids were recorded in the middle Villafranchian and were associated with open and dry habitats (steppes), whereas the small ones lived in more closed and wet habitats (savanna-mosaic, woodlands) from late Villafranchian to earliest Galerian.

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Research Article
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Copyright © The Paleontological Society 

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