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Lichidae (Trilobita): Morphology and classification

Published online by Cambridge University Press:  14 July 2015

H. B. Whittington*
Affiliation:
Department of Earth Sciences, University of Cambridge, Downing Street, Cambridge CB2 3EQ, United Kingdom

Abstract

Morphology is reviewed with special reference to the doublure and thoracic characters; a reconstruction of Richterarges aquilonius is used to suggest possible lichid anatomy. The relatively large hypostome and wide doublure, and the thorax with, or without, the strongly convex posterior pleural bands characterize lichids, in addition to the distinctive glabellar morphology. Relatively shallow furrows on the external surface form strong ridges on the visceral surface of the exoskeleton; apodemes are absent. Type specimens of species described by Schmidt, recently traced in Russian museums, are figured. Lichids differ from odontopleurids in morphology and anatomy, and are unlike Scutelluinae; accepted subfamilial divisions of Lichidae are reinforced by thoracic characters. A single lichid species is known in the early Tremadoc, in the early Middle Ordovician the main four subfamilies are recognised and are widespread; their ancestry and early diversification remain unknown.

Type
Research Article
Copyright
Copyright © The Paleontological Society

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References

Adrain, J. M. 1994. The lichid trilobite Borealarges n. gen., with species from the Silurian of Arctic Canada. Journal of Paleontology, 68:10811099.CrossRefGoogle Scholar
Adrain, J. M., and Ramskold, L. 1996. The lichid trilobite Radiolichas in the Silurian of Arctic Canada and Gotland, Sweden. Geological Magazine, 133:147158.CrossRefGoogle Scholar
Barrande, J. 1846. Notice préliminaire sur le Systême Silurien et les trilobites de Bohême. Hirschfeld, Leipzig, 96 p.CrossRefGoogle Scholar
Barrande, J. 1852. Systême Silurien du centre de la Bohême. Ière Partie: Recherches Paléontologiques. Volume 1. Prague and Paris, 935 p.Google Scholar
Barrande, J. 1872. Systême Silurien du centre de la Bohême. Ière Partie: Recherches Paléontologiques, Volume 1, Supplement. Prague and Paris, 647 p.Google Scholar
Basse, M. 1998. Trilobiten aus mittlerem Devons des Rhenohercynikums: III. Palaeontographica, Abteilung A, 249:1162.Google Scholar
Beecher, C. E. 1897. Outline of a natural classification of the trilobites. American Journal of Science, Series 4, 3:89106, 181–207.CrossRefGoogle Scholar
Bergström, J. 1973. Organization, life and systematics of trilobites. Fossils and Strata, 2, 69 p.Google Scholar
Beyrich, E. 1845. Ueber einige böhmischer Trilobiten. Reimer, Berlin, 47 p.Google Scholar
Bigsby, J. J. 1825. Description of a new species of trilobite. Journal of the Academy of Natural Science, Philadelphia, 4:365368.Google Scholar
Billings, E. 1861–1865. Palaeozoic Fossils. Volume 1. Geological Survey of Canada, 426 p.Google Scholar
Brett, C. E., and Taylor, W. L. 1997. The Homocrinus Beds: Silurian crinoid lagerstätten of western New York and southern Ontario, p. 181223. In Brett, C. E. and Baird, G. C. (eds.), Paleontological Events. Stratigraphic, Ecological and Evolutionary Implications. Columbia University Press, New York, xv + 604 p.Google Scholar
Brett, C. E., Whiteley, T. E., Allison, P. A., and Yochelson, E. L. 1999. The Walcott-Rust quarry: Middle Ordovician trilobite Konservat-Lagerstätten. Journal of Paleontology, 73:288305.CrossRefGoogle Scholar
Bruton, D. L. 1966. A revision of the Swedish Ordovician Odontopleuridae (Trilobita). The Bulletin of the Geological Institutions of the University of Uppsala, 43:140.Google Scholar
Bruton, D. L., and Haas, W. 1997. Functional morphology of Phacopinae (Trilobita) and the mechanics of enrolment. Palaeontographica, Abteilung A, 245:143.Google Scholar
Bruton, D. L., and Haas, W. 1999. The anatomy and functional morphology of Phacops (Trilobita) from the Hunsrück Slate (Devonian). Palaeontographica, Abteilung A, 253:2975.Google Scholar
Bruton, D. L., Hoel, O. A., Beyenne, L. T., and Ivantsov, A. Yu. 1997. Catalogue of the trilobites figured in Friedrich Schmidt's “Revision der ostbaltischen silurischen Trilobiten” (1881–1907). Contributions from the Palaeontological Museum, University of Oslo, 403, 117 p.Google Scholar
Castelnau, F. de. 1843. Essai sur le système silurien de l'Amérique Septentrionale. Paris and Strasbourg, 56 p.CrossRefGoogle Scholar
Chatterton, B. D. E. 1971. Taxonomy and ontogeny of Siluro-Devonian trilobites from near Yass, New South Wales. Palaeontographica, Abteilung A, 137:1108.Google Scholar
Chatterton, B. D. E. 1980. Ontogenetic studies of Middle Ordovician trilobites from the Esbataottine Formation, MacKenzie Mountains, Canada. Palaeontographica, Abteilung A, 171:174.Google Scholar
Chatterton, B. D. E., and Campbell, M. 1993. Enrolling in trilobites: a review and some new characters. Memoirs of the Association of Australasian Palaeontologists, 15:103123.Google Scholar
Chatterton, B. D. E., and Ludvigsen, R. 1976. Silicified Middle Ordovician trilobites from the South Nahanni River area, District of Mackenzie, Canada. Palaeontographica, Abteilung A, 154:1106.Google Scholar
Chatterton, B. D. E., Johnson, B. D., and Campbell, K. S. W. 1979. Silicified Lower Devonian trilobites from New South Wales. Palaeontology, 22:799837.Google Scholar
Chlupáč, I. 1983. Trilobite assemblages in the Devonian of the Barrandian area and their relations to palaeoenvironments. Geologica et Palaeontologica, 17:4573.Google Scholar
Chlupáč, I. 1987. Ecostratigraphy of Silurian trilobite assemblages of the Barrandian area, Czechoslovakia. Newsletters on Stratigraphy, 17(3):169186.CrossRefGoogle Scholar
Chlupáč, I. 1999. Barrande's stratigraphic concepts, palaeontological localities and tradition—comparison with the present state. Journal of the Czech Geological Society, 44:330.Google Scholar
Cocks, L. R. M. 2000. The early Palaeozoic Geography of Europe. Journal of the Geological Society, London, 157:110.CrossRefGoogle Scholar
Eichwald, E. D. 1859–1861. Lethea Rossica, ou paléontologie de la Russie, Volume 1 (Animaux fossiles de la période ancienne), pt. 2, p. 2711633. E. Schweizerbart, Stuttgart. Volume 1 of this work was issued in three parts (see review in Quarterly Journal of the Geological Society, London, 1862, Volume 18, pt. 2, miscellaneous, p. 3–4), the section on trilobites being on p. 1379–1390, published in 1860, 7th livraison, p. 1005–1532.Google Scholar
Feist, R. 1991. The late Devonian crises. Historical Biology, 5:197214.CrossRefGoogle Scholar
Fletcher, T. W. 1850. Observations on Dudley trilobites. Quarterly Journal of the Geological Society of London, 6:235239.CrossRefGoogle Scholar
Fortey, R. A. 1990. Ontogeny, hypostome attachment and trilobite classification. Palaeontology, 33:529576.Google Scholar
Fortey, R. A., and Owens, R. M. 1999. Feeding habits in trilobites. Palaeontology, 42:429465.CrossRefGoogle Scholar
Gürich, G. 1901. Ueber ein neue Lichas-Art aus dem Devon von Neu-Süd Wales und über die Gattung Lichas überhaupt. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, 14:519539.Google Scholar
Hawle, I., and Corda, A. J. C. 1847. Prodrom einer Monographic der Böhmischen Trilobiten. Abhandlungen Koeniglichen Boehmischer Gesellschaft der Wissenschaften. Prague, 176 p.Google Scholar
Henningsmoen, G. 1951. Remarks on the classification of trilobites. Norsk geologisk tiddskrift, 29:174217.Google Scholar
Horný, R., and Bastl, F. 1970. Type specimens of fossils in the National Museum, Prague. Volume 1. Trilobita. National Museum, Prague, 354 p.Google Scholar
Hupé, P. 1953. Classe des Trilobites, p. 44246. In Piveteau, J. (ed.), Traité de Paléontologie. Volume 3. Masson, Paris.Google Scholar
Hupé, P. 1955. Classification des Trilobites. Annales de Paléontologie, 41:91325.Google Scholar
Kaesler, R. L. (ed.). 1997. Treatise on Invertebrate Paleontology, Pt. O, Arthropoda 1, Trilobita, revised. Volume 1. Geological Society of America and University of Kansas Press, Lawrence, 530 p.Google Scholar
Kowalski, H. 1992. Trilobiten, verwandlung-künstler der Paläozoikums. Korb, Goldschneck-Verlag Weidert, 157 p.Google Scholar
Lamont, A. 1969. Prolegomena to aggressive mimicry and protective resemblance in early fishes, chelicerates, trilobites and brachiopods. Scottish Journal of Science, 1:75103.Google Scholar
Lindström, G. 1901. Researches on the visual organs of the trilobites. Kong Svenska Vetenskaps-Akademiens Handlingar, 34(8), 86 p.Google Scholar
Ludvigsen, R. 1979. Fossils of Ontario, Pt. 1, The trilobites. Life Sciences Miscellaneous Publications, Royal Ontario Museum, Toronto, 96 p.Google Scholar
Mergl, M. 1994. Trilobite fauna from the Trěnice Formation (Tremadoc) in central Bohemia. Folia Musei Rerum naturalium Bohemiae Occidentalis, Geologica 39, 31 p.Google Scholar
Milne Edwards, H. 1840. Histoire naturelle des Crustacés, comprenant l'anatomie, le physiologie et la classification de ces animaux, Volume 3, Paris, 638 p.Google Scholar
Moore, R. C. 1959. Treatise on Invertebrate Paleontology, Pt. O, Arthropoda 1. Geological Society of America and University of Kansas Press, Lawrence, 560 p.Google Scholar
Morris, S. F., and Fortey, R. A. 1985. Catalogue of the type and figured specimens of Trilobita in the British Museum (Natural History). British Museum (Natural History), London, 183 p.Google Scholar
Phleger, F. B. 1936. Lichadian trilobites. Journal of Paleontology, 10:593615.Google Scholar
Phleger, F. B. 1937. New Lichadacea in the collections of the Museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology, Harvard College, 80:415423.Google Scholar
Přibyl, A., and Erben, H. K. 1952. Über einige neue oder wenig bekannte Acanthopyginae (Tril.) des böhmischen und des deutschen Devons. Paláontologia Zeitschrift, 26:141174.CrossRefGoogle Scholar
Rabano, I. 1989. El género Uralichas Delgado, 1892 (Trilobita, Lichaida) en el Ordovícico de la Península Ibérica. Boletin Geológico y Minero, 100:2147.Google Scholar
Ramsköld, L. 1991a. The perforated trilobite Laethoprusia gen. nov. and the phylogeny of Koneprusia and Isoprusia (Odontopleuridae, Koneprusiinae). Transactions of the Royal Society of Edinburgh, Earth Sciences, 82:125141.CrossRefGoogle Scholar
Ramsköld, L. 1991b. Pattern and process in the evolution of the Odontopleuridae (Trilobita). The Selenopeltinae and Ceratocephalinae. Transactions of the Royal Society of Edinburgh, Earth Sciences, 82:143181.CrossRefGoogle Scholar
Raymond, P. E. 1905. Note on the names Amphion, Harpina, and Platymetopus . American Journal of Science, 19:377378.CrossRefGoogle Scholar
Reed, F. R. C. 1902. Notes of the genus Lichas . Quarterly Journal of the Geological Society of London, 58:5982.CrossRefGoogle Scholar
Reed, F. R. C. 1907. A new species of Lichas . Geological Magazine, New Series, Decade 5, Volume 4:396400.CrossRefGoogle Scholar
Reed, F. R. C. 1914. The Lower Palaeozoic trilobites of Girvan. Supplement. Palaeontographical Society, London, 56 p.Google Scholar
Reimann, I. G. 1941. A new restoration of Terataspis . Bulletin of the Buffalo Society of Natural Sciences, 17:3946.Google Scholar
Reimann, I. G. 1943. Terataspis gets remodeling. Hobbies, Buffalo Museum of Science, 23:7577.Google Scholar
Reimann, I. G. 1945. The thorax of Terataspis . Journal of Paleontology, 19:6991.Google Scholar
Richter, R., and Richter, E. 1917. Die Lichadiden des Eifler Devons. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, Jahrgang 1917, Pt. 1:572.Google Scholar
Růžička, R. 1926. Faune des couches à Euloma de gisement ferrugineaux près de Holubkuov (à Ouzky). Bulletin international de l'Acadèmie des Sciences de Bohême, 24 p.Google Scholar
Růžička, R. 1935. Příspěvek k poznaí trilobitů Barrandienu. Rozpravy 2. Třidy Ceské Akademie, 44, no. 37, 8 p.Google Scholar
Šnajdr, M. 1990. Bohemian trilobites. Published by the Geological Survey, Prague, 265 p.Google Scholar
Schmidt, F. 1885. Revision der ostbaltischen silurischen trilobiten, Abteilung 2, Acidaspiden und Lichiden. Mémoires de l'Académie Impériale des Sciences de St. Pétersbourg, 7th Series, 33(1):127 p.Google Scholar
Thomas, A. T. 1979. Trilobite associations in the British Wenlock, p. 447451. In Harris, A. L., Holland, C. H., and Leake, B. E. (eds.), The Caledonides of the British Isles—reviewed. Geological Society of London, Scottish Academic Press.Google Scholar
Thomas, A. T. 1981. British Wenlock trilobites, Pt. 2, Palaeontographical Society Monographs, 134:5799.Google Scholar
Thomas, A. T., and Holloway, D. J. 1988. Classification and phylogeny of the trilobite Order Lichida. Philosophical Transactions of the Royal Society of London, 321:179262.Google Scholar
Thorsteinsson, R., and Uyeno, T. T. 1980. Stratigraphy and conodonts of Upper Silurian and Lower Devonian rocks in the environs of the Boothia Uplift, Canadian Arctic Archipelago. Geological Survey of Canada, Bulletin, 292:174.Google Scholar
Tripp, R. P. 1957. The classification and evolution of the Superfamily Lichacea. Geological Magazine, 94:104122.CrossRefGoogle Scholar
Tripp, R. P. 1958. Stratigraphical and geographical distribution of the named species of the trilobite Superfamily Lichacea. Journal of Paleontology, 32:574582.Google Scholar
Vaněk, J. 1959. Čeled Lichaidae Hawle et Corda, 1847 ze středočeského staršího paleozoika (Trilobitae). Bohemia Centralis (A—Scientiae naturales), 1(3):81168.Google Scholar
Vaněk, J. 1990. Exact age of several localities in the Liten Formation, Motol Member (Wenlock, Central Bohemia). Věstník Ustředního ústavu geologického, 65:119124.Google Scholar
Vaněk, J. 1999. On the genus Dicranogmus Hawle and Corda, 1847 (Trilobita, Lichidae). Palaeontologia Bohemiae, 5:3335.Google Scholar
Warburg, E. 1925. The trilobites of the Leptaena Limestone in Dalarne. Bulletin of the Geological Institute of Upsala, 17:viii + 446 p.Google Scholar
Whittington, H. B. 1956. Silicified Middle Ordovician trilobites: the Odontopleuridae. Bulletin of the Museum of Comparative Zoology at Harvard College, 114:155288.Google Scholar
Whittington, H. B. 1961. Silurian Hemiarges (Trilobita) from Cornwallis Island and New York State. Journal of Paleontology, 35:433444.Google Scholar
Whittington, H. B. 1963. Middle Ordovician trilobites from Lower Head, western Newfoundland. Bulletin of the Museum of Comparative Zoology at Harvard College, 129:1118.Google Scholar
Whittington, H. B. 1988. Hypostomes of post-Cambrian trilobites. New Mexico Bureau of Mines and Mineral Resources, Memoir 44:321339.Google Scholar
Whittington, H. B. 1992. Trilobites. Boydell and Brewer, Woodbridge, U.K. and Rochester, N.Y., 145 p., 120 pls.Google Scholar
Whittington, H. B. 1993a. Anatomy of the Ordovician trilobite Placoparia . Philosophical Transactions of the Royal Society, London, B 339:109118.Google Scholar
Whittington, H. B. 1993b. Morphology, anatomy and habits of the Silurian homalonotid trilobite Trimerus . Memoirs of the Association of Australasian Palaeontologists, 15:6983.Google Scholar
Whittington, H. B. 1999. Siluro-Devonian Scutelluinae (Trilobita) from the Czech Republic: morphology and classification. Journal of Paleontology, 73:414430.CrossRefGoogle Scholar
Whittington, H. B., and Almond, J. E. 1987. Appendages and habits of the Upper Ordovician trilobite Triarthus eatoni . Philosophical Transactions of the Royal Society, London, B 317:146.Google Scholar