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New and revised cyrtospiriferid (Spiriferida) brachiopods from the lower Famennian (Upper Devonian) of Armenia

Published online by Cambridge University Press:  31 March 2022

Vahram Serobyan
Affiliation:
Institute of Geological Sciences of the National Academy of Sciences of the Republic of Armenia-Yerevan, Armenia, 24A, Marshal Baghramyan Avenue, Yerevan 0019, Republic of Armenia , University of Lille, CNRS, UMR 8198- Evo-Eco-Paleo, F-59000 Lille, France , ,
Taniel Danelian*
Affiliation:
University of Lille, CNRS, UMR 8198- Evo-Eco-Paleo, F-59000 Lille, France , ,
Catherine Crônier
Affiliation:
University of Lille, CNRS, UMR 8198- Evo-Eco-Paleo, F-59000 Lille, France , ,
Araik Grigoryan
Affiliation:
Institute of Geological Sciences of the National Academy of Sciences of the Republic of Armenia-Yerevan, Armenia, 24A, Marshal Baghramyan Avenue, Yerevan 0019, Republic of Armenia ,
Bernard Mottequin
Affiliation:
Operational Directorate Earth and History of Life, Royal Belgian Institute of Natural Sciences, rue Vautier 29, B 1000 Brussels, Belgium
*
*Corresponding author.

Abstract

The Upper Devonian sedimentary sequences of Central Armenia, which mainly consist of shallow water, mixed carbonate-siliciclastic deposits, contain abundant and diverse brachiopods that are dominated by spiriferides. Based on newly collected material from the lower Famennian Aramazdospirifer orbelianus brachiopod zone (coeval to the Palmatolepis crepida conodont zone) of Armenia, we here introduce two new cyrtospiriferid genera and fully document their type species, including their intraspecific variability. Pentagonospirifer n. gen. is a monospecific genus assigned to the subfamily Cyrtospiriferinae that is currently known only from the lower Famennian of Armenia; it is likely that its type species, P. abrahamyanae n. sp., evolved from the species Cyrtospirifer verneuili sensu Abrahamyan, known from the Frasnian and lower Famennian of Armenia. The second new genus, Tornatospirifer n. gen., is assigned to the subfamily Cyrtiopsinae and defined on the basis of one of the most biostratigraphically valuable cyrtospiriferid species, namely T. armenicus n. comb., described previously from the lower Famennian Aramazdospirifer orbelianus Zone of Armenia. It is likely that it evolved from a Frasnian ancestral stock of Tiocyrspis Sartenaer, known from northwestern Europe, which probably migrated during the early Frasnian to the north Gondwanan margin. A neotype is selected for Abrahamyan's species because the type material is lost. Epibionts (Cornulites, Hederella) attached to brachiopods are also documented for the first time from the Upper Devonian of Armenia.

UUID: http://zoobank.org/923d232f-cd6a-48bb-8a05-27c689058b65.

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Copyright © The Author(s), 2022. Published by Cambridge University Press on behalf of The Paleontological Society

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