We consider a dynamic multilevel population or information system. At each level individuals or information units undergo a Galton–Watson-type branching process in which they can be replicated or removed. In addition, a collection of individuals or information units at a given level constitutes an information unit at the next higher level. Each collection of units also undergoes a Galton–Watson branching process, either dying or replicating. In this paper, we represent this multilevel branching model as a measure-valued stochastic process, study its moment structure, identify the limiting continuous-state approximation and analyse the long-time behavior in both non-critical and critical cases. For example, we obtain an asymptotic expression for the extinction probability for the total population mass process and an analogue of Yaglom's conditioned limit theorem in the critical case.
