In the Central Iberian Zone, the Cadomian orogenic collapse is represented by chaotic megabreccias, olistostromes and mélange deposits reflecting a drastic change from slope-related deposits, fed by denudation of the Cadomian arc, to offshore-dominant settings episodically punctuated by phosphogenetic processes. In the Ibor and Alcudia anticlines, the pre-rift unconformity is marked by paraconformable to angular discordant contacts separating variable tilted strata of the Ediacaran Lower Alcudian – Domo Extremeño Supergroup and the upper Ediacaran – lower Terreneuvian Ibor Group from the overlying Terreneuvian San Lorenzo and Fuentepizarra formations. The sedimentation of the San Lorenzo Formation reflects two palaeogeographic scenarios: (i) a low-angle stable basement recording shoaling-upward siliciclastic cycles, and (ii) perturbations of basement fault scarps feeding slope-related conglomeratic channels, with NE-directed palaeocurrents, and sourced from topographic palaeohighs controlled by the movement along synsedimentary normal fault systems, such as the so-called El Guijo Fault. The intra-Fortunian age of the pre-rift unconformity is constrained by the ichno- and microfossil content of the succession, and is bracketed between the first occurrence of Treptichnus pedum in the Arrocampo Formation (Ibor Group) and of Anabarella plana in the Fuentepizarra Formation.