For more than fifty years it has been known that mammalian faunas of late-Pleistocene age are taxonomically unique and lack modern analogs. It has long been thought that nonanalog mammalian faunas are limited in North America to areas east of the Rocky Mountains and that late-Pleistocene mammalian faunas in the west were modern in taxonomic composition. A late-Pleistocene fauna from Marmes Rockshelter in southeastern Washington State has no modern analog and defines an area of maximum sympatry that indicates significantly cooler summers than are found in the area today. An earliest Holocene fauna from Marmes Rockshelter defines an area of maximum sympatry, including the site area, but contains a single tentatively identified taxon that may indicate slightly cooler than modern summers.
