January Cave, in the Rocky Mountains of southwestern Alberta, has yielded vertebrate remains from a coprocenosis of mid-Wisconsinan-age. Taphonomic analysis indicates accumulation by raptors, mostly owls, and mammalian carnivores. The vertebrate record, together with pollen analysis, indicates that cool, dry conditions prevailed in an extensive tundra-like environment, with prairie elements in the valleys below. Thirty-four mammalian taxa have been recovered from January Cave. Today, some of these species (e.g., Lemmus sibiricus and Dicrostonyx torquatus) do not coexist with others (e.g., Cynomys sp., Mustela nigripes, Vulpes velox, and Lagurus curtatus). Therefore, the January Cave local fauna represents a “nonanalog” mammalian community characteristic of the late Pleistocene. It suggests that the region enjoyed an equable climate, with reduced climatic extremes but still cool, further supporting a mid-Wisconsinan age estimate for the fauna. It is the first major, small vertebrate fauna of its age to be reported from Alberta.