The late Brigantian topmost parts of the Pathhead Formation (Aberlady Formation in East Lothian) and the succeeding Lower Limestone Formation crop out widely in Fife and East Lothian. The successions include nine deltaic, coastal floodplain and marine shelf cycles (cyclothems), of which the lowest examined terminates the Pathhead and Aberlady Formations and the remaining eight constitute the Lower Limestone Formation.

The cyclothems conform broadly to the ‘Yoredale” transgressive/regressive pattern in which a transgressive marine shelf phase is succeeded by delta progradation and terminates with a fluvial delta plain phase. Cycles may combine to form compound cyclothems up to more than 50 m thick, in which a basal, typically complete initial cycle of Yoredale pattern is succeeded by up to five base-absent minor cycles. These are thinner, more variable and less laterally persistent units in which the marine phase is weakly represented or absent.

Cyclothems reflect successive marine flooding events, possibly under eustatic control, succeeded by delta progradation and, ultimately, leading to extensive palaeosol formation, including coal seams. Sedimentation and palaeosol formation were partly controlled by fault-induced differential subsidence and are likely to have been related to autocyclic processes. Local uplift and subsidence associated with vulcanicity, as at Kinghorn and Elie, have led to thickening or thinning of sediments accumulated in a given time period.

Initial cycles initiate longer-period allocycles, corresponding broadly to third-order Exxon Production & Research (EPR) Type 1 sequences having a periodicity of around 1 Ma, within the Milankovitch orbital band. Two parasequences constitute each initial cycle: a lower, initiated on a marine flooding surface, and an upper, bounded by the base of the lowest thick sandstone in the cycle; cyclothem bases and sequence bases thus alternate. Parasequences and sequences are less well defined in minor cycles due to the problem of tracing the combined disconformity and soil profile of the underclay beyond the edge of channel sandstones. Minor cycles were controlled primarily by short-period autocyclic sedimentary and, or, tectonic processes, including delta-lobe switching and differential subsidence.

Although we have attempted to interpret the deposits of Fife and the Lothians in terms of sequence stratigraphy, we are not fully convinced that the patterns of associated changes widely recognised within the framework of sequence stratigraphy can be confidently applied in succesions in which autocyclic changes feature strongly in an area undergoing active basin subsidence associated with strike-slip faulting. There is no doubt that some of the cyclicity discerned in the late Brigantian successions of eastern Scotland was related to eustatic sea level changes, which gave rise to the widespread limestone platforms or marine bands. The formation of eight cyclothems within the 2·5–3·5 Ma of late Brigantian suggests a cyclicity of about 400 ka, which corresponds to the long period eccentricity cycles of Milankovitch rather than the 0·5–5·0 Ma of third-order EPR cycles.

This paper completes the description of intact and three-dimensional Ctenopyge species from the upper Cambrian Peltura minor Zone in Västergötland, central Sweden. All these species are present together, on the same bedding planes. The most abundant species, Ctenopyge (Eoctenopyge) angusta Westergård, 1922 has previously been described, and an almost complete ontogeny worked out. C. (Ctenopyge) gracilis Henningsmoen, 1957 is a small trilobite with nine thoracic segments and very long, thin curving and subparallel thoracic spines; the genal spines partially encircle the body. Two axial spines at the rear are of considerable length. When reconstructed in side view, the posterior thoracic spines rise upwards as an inclined fan, but when relaxed the tips of all the thoracic and axial spines come to lie in the same plane as the horizontal genal spines. An almost complete ontogeny is described for this species, and individuals show an evident spinosity from an early stage, but the body size at which thoracic segments are liberated is highly variable. C. (Ctenopyge) ahlbergi n. sp. is a larger, robust and broad species distinguished by long, stout genal spines, ten thoracic segments, and a very spiny body with the first three to four spines expanded into lateral flanges. A degree 6 meraspis shows these flanges already developing. C. (Ctenopyge) rushtoni n. sp has likewise ten thoracic segments, and has stout, broad-based and tapering spines. Incomplete meraspides 6 and 7 are known for this species. In both C. (Ctenopyge) ahlbergi and C. (Ctenopyge) rushtoni there are also two axial spines at the rear, and the extended body would have had a similar rising tail fan to that of C. (Ctenopyge) gracilis. C. (Mesoctenopyge) tumida is also present as a single large adult and several smaller holaspides. In this species the first thoracic segment is confirmed as bearing a pair of long curving spines, somewhat smaller than the encircling genal spines. The remaining thoracic spines are straight and sharp, and evidently longer in young holaspides. There is a single long axial spine on the last segment. No adult pygidium has been found.

Some comments on the diversity of the fauna as a whole and the range of functional types are appended.

The dental morphology of the pycnodontid fish †Stemmatodus rhombus (Agassiz, 1844) from the Lower Cretaceous (lower Barremian) of southern Italy is described in detail. SEM pictures of vomerine and prearticular teeth are provided for the first time allowing the comparison with other Early Cretaceous pycnodonts, which are only known from isolated dental remains. The dentition of †Stemmatodus rhombus consists of incisiform grasping teeth on the paired premaxillae and dentaries and crushing teeth on the unpaired vomer and paired prearticulars. The vomerine teeth are arranged in five longitudinal rows, which display symmetry with a distinct median principal and two lateral rows. The prearticular bears three longitudinal rows. Teeth of the main row are much wider than those of the lateral ones. The tooth crowns are rather flat and expose a shallow apical depression, which is surrounded by a ring of widely-spaced small tubercles. The ornamentation and arrangement of teeth distinguishes dentitions of †Stemmatodus from all other pycnodonts. The wear pattern of the teeth is evaluated and the significance of the dental morphology in pycnodont taxonomy is discussed. In addition, the systematic position of †Stemmatodus is discussed.

The recently published model of evolution and biogeography and the resulting classification of the Tetraconodontinae (Van der Made 1999) were criticised by Pickford (2001) on eight points. It was claimed that Conohyus giganteus is a suine, and that this ‘radically modifies tetraconodont systematics’. However, Pickford failed to demonstrate that the type specimen is a suine, and even if it were a suine, this would affect only one of over twenty species names; not a radical change of the proposed evolutionary model.