The Oligocene strata of the Qom Formation from the Tajar-Kuh section, Central Iran, are rich in various Larger Benthic Foraminifera (LBF). Morphometric studies of the internal features of the LBF tests were carried out based on matrix-free specimens from seven samples. The LBF assemblage studied in 24 populations mainly contains representatives of two nummulitid (the reticulate Nummulites bormidiensis Tellini, 1888, only in the lower three samples, and the radiate Nummulites kecskemetii Less, 1991 in all samples) and of two lepidocyclinid lineages. Of the latter, Nephrolepidina praemarginata (R. Douvillé, 1908) occurs in all the samples, whereas Eulepidina formosoides H. Douvillé, 1925 is present in the lower six samples. In the upper sample, the slightly more advanced Eulepidina ex. interc. formosoides H. Douvillé, 1925 et dilatata (Michelotti, 1861) is recorded. Due to the very similar taxonomic composition of the Tajar-Kuh section with their coeval faunas of the Mediterranean, the age was evaluated in the frame of the Western Tethyan Oligo-Miocene shallow benthic zonation (SBZ). Although the presence of E. formosoides suggests late Rupelian (SBZ 22A Zone) age, the occurrence of Heterostegina assilinoides, N. kecskemetii, N. bormidiensis, and Planolinderina sp. preferably represents the SBZ 22B Zone of the early Chattian. Based on the obtained results, at least the lower six samples can indicate the very basal part of the Chattian. More advanced Eulepidina from the uppermost sample suggests a slightly younger but still early Chattian age.

The recent description of the nevadioid trilobite Buenellus chilhoweensis Webster and Hageman, 2018 established the presence of early Cambrian Montezuman Stage (Cambrian Series 2, Stage 3) faunas in the Murray Shale of Chilhowee Mountain, Tennessee. The description recognized the oldest known age-diagnostic Cambrian trilobite from the Laurentian margin of the former Iapetus Ocean since Buenellus Blaker, 1988 is known otherwise only from the Sirius Passet Lagerstätte on the Innuitian margin of North Greenland. The bivalved arthropods Isoxys chilhoweanus Walcott, 1890 and Indota tennesseensis (Resser, 1938a) have also been described from the Murray Shale, but hyolithids appear to be the dominant body fossils in terms of diversity and abundance. Although poorly preserved, the hyolithids occurring together with Buenellus chilhoweensis are described to improve understanding of the Murray Shale biota. The hyolith assemblages of the Murray Shale and Sirius Passet Lagerstätte are not closely similar, although the poor preservation of both hinders comparison.

In recent years the plethora of ‘weird wonders,’ the vernacular for the apparently extinct major body plans documented in many of the Cambrian Lagerstätten, has been dramatically trimmed. This is because various taxa have been either assigned to known phyla or accommodated in larger monophyletic assemblages. Nevertheless, a number of Cambrian taxa retain their enigmatic status. To this intriguing roster we add Dakorhachis thambus n. gen. n. sp. from the Miaolingian (Guzhangian) Weeks Formation Konservat-Lagerstätte of Utah. Specimens consist of an elongate body that lacks appendages but is apparently segmented. A prominent feeding apparatus consists of a circlet of triangular teeth, while posteriorly there are three distinct skeletal components. D. thambus is interpreted as an ambush predator and may have been partially infaunal. The wider affinities of this new taxon remain conjectural, but it is suggested that it may represent a stem-group member of the Gnathifera, today represented by the gnathostomulids, micrognathozoans, and rotifers and possibly with links to the chaetognaths.

UUID: http://zoobank.org/22113e6b-f79e-4d06-9483-144618a61327

A Late Ordovician brachiopod fauna from the Black River quadrangle (D-1 1:63,360 scale) of east-central Alaska comprises taxa typical of the Late Ordovician brachiopod fauna in the pericratonic epeiric seas of Laurentia, including Hesperorthis pyramidalis (Twenhofel, 1928), Plaesiomys occidentalis (Okulitch, 1943), Eoplectodonta sp., Holtehdalina sp., Leptaena sp., Brevilamnulella minuta n. sp., Tcherskidium tenuicostatum n. sp., Rhynchotrema iowense Wang, 1949, and Whitfieldella sp. The presence of Plaesiomys occidentalis and Tcherskidium tenuicostata n. sp. indicates a latest Katian age by correlation with similar species in the Mackenzie Mountains, southern Manitoba, Anticosti Island, the American midcontinent, Kolyma, and Siberia. Cluster analysis based on 20 well-studied late Katian brachiopod faunas from various regions within Laurentia and elsewhere in other tectonic plates suggests that the small brachiopod faunule from Alaska has the strongest paleobiogeographic affinity with Laurentia, confirming that the Black River quadrangle of Alaska was part of Laurentia during the Late Ordovician.

UUID: http://zoobank.org/6b387856-61d5-4685-a592-faf5287c1e0f

A well-preserved fauna of largely articulated trilobites is described from three new localities close to one another in the Bull River Valley, southeastern British Columbia. All the trilobites from these localities are from the lower or middle part of the Wujiajiania lyndasmithae Subzone of the Elvinia Zone, lower Jiangshanian, in the McKay Group. Two new species are proposed with types from these localities: Aciculolenus askewi and Cliffia nicoleae. The trilobite (and agnostid) fauna from these localities includes at least 20 species: Aciculolenus askewi n. sp., Agnostotes orientalis (Kobayashi, 1935), Cernuolimbus ludvigseni Chatterton and Gibb, 2016, Cliffia nicoleae n. sp., Elvinia roemeri (Shumard, 1861), Grandagnostus? species 1 of Chatterton and Gibb, 2016, Eugonocare? phillipi Chatterton and Gibb, 2016, Eugonocare? sp. A, Housia vacuna (Walcott, 1912), Irvingella convexa (Kobayashi, 1935), Irvingella flohri Resser, 1942, Irvingella species B Chatterton and Gibb, 2016, Olenaspella chrisnewi Chatterton and Gibb, 2016, Proceratopyge canadensis (Chatterton and Ludvigsen, 1998), Proceratopyge rectispinata (Troedsson, 1937), Pseudagnostus cf. P. josepha (Hall, 1863), Pseudagnostus securiger (Lake, 1906), Pseudeugonocare bispinatum (Kobayashi, 1962), Pterocephalia sp., and Wujiajiania lyndasmithae Chatterton and Gibb, 2016. Pseudagnostus securiger, a widespread early Jiangshanian species, has not been previously recorded from southeastern British Columbia. Non-trilobite fossils collected from these localities include brachiopods, rare trace fossils, a complete silica sponge (Hyalospongea), and a dendroid graptolite. The faunas from these localities are more diverse and better preserved than those from other previously documented localities of the same age in the region.

Additional specimens of a rare species, found by amateur collectors in previously documented localities of slightly younger age (upper part of Wujiajiania lyndasmithae Subzone) in the same region, are documented. These new specimens, when combined with an earlier discovered specimen, provide adequate type material to propose a new species of Labiostria, L. gibbae, which may be useful for biostratigraphy.

UUID:http://zoobank.org/89551eac-b3af-4b2b-8ef3-7c2e106a560d

The dalmanitid trilobite Kasachstania Maksimova, 1972, previously reported from the Lower Devonian of Kazakhstan and North America (USA) and the upper Silurian–Lower Devonian of South America (Bolivia and Argentina), is revised. Kasachstania kasachstanica (Balashova in Maksimova,1968) and K. septicostata (Maksimova, 1968) are regarded as junior synonyms of the type species K. saryarkensis (Maksimova, 1960), all from the Lower Devonian of the type locality in central Kazakhstan (northern Balkhash). On the basis of a new diagnosis, K. ulrichi ulrichi (Delo, 1940) from the Emsian of the United States, K. ulrichi asiatica (Maksimova, 1968), K. pristina (Maksimova, 1968), and K. alperovichi Pour et al., 2019, from the Lower Devonian of Kazakhstan, K. andii (Kozłowski, 1923) from the upper Silurian–Lower Devonian of Bolivia, and K. gerardoi Edgecombe and Ramsköld, 1994, from the upper Silurian–Lower Devonian of Bolivia and Argentina are excluded from Kasachstania. This genus, represented only by K. saryarkensis and K. kiikbaica (Maksimova, 1968), is restricted to the Lower Devonian of central Kazakhstan, corresponding to the Balkhash–Mongolo–Okhotsk province in the paleobiogeographic context of the Old World Realm, instead of being nearly cosmopolitan as previously considered. In addition, we provide some remarks about Saryarkella Maksimova, 1978b, a monospecific dalmanitid genus largely overlooked although valid from the Emsian of the same area in central Kazakhstan.

The fossil record of the family Camerobiidae has been represented by only one species, Neophyllobius succineus Bolland and Magowski, 1990, described from Eocene Baltic amber. These prostigmatan mites are distinguishable by their distinctly long and slender stilt-like legs, and they are associated with aboveground vegetation where they hunt for other small invertebrates. This paper enhances the knowledge of fossil stilt-legged mites. Two new fossil species, N. electrus new species and N. glaesus new species, are described from samples of Baltic amber, and remarks on their morphology and taphonomy are provided. The discovery is complemented with a discussion on morphological singularities (the shape of the prodorsum, the location of setae h1 and h2 in living specimens, and lengths of genual setae), an anomaly of hypertrophied seta (found in the N. glaesus holotype), and some biogeographical issues.

UUID: http://zoobank.org/d1602384-ae4f-4f90-b4a1-6cdedd77c9e1

Two new isocrinids are described from the Lower Cretaceous Agrio Formation of the Neuquén Basin, west-central Argentina. Isocrinus (Chladocrinus) covuncoensis new species is based on several beautifully preserved specimens from Valanginian beds of the Pilmatué Member. It is characterized by a small size, multiramose crown with six arm divisions, 240 arm tips, mostly isotomous branching, seven (or rarely eight) secundibrachials, smooth and stout column, short noditaxis, and pentalobate columnals. The species occurs in a 30 m thick interval of cross-bedded sandstones and mixed clastic-carbonate sediments that represent the migration of large, tidally influenced, subaqueous dunes developed in the offshore. Sudden burial of crinoids that dwelled on the dune toes and interdunes, possibly by the acceleration of the lee face migration, provided the exceptional preservation of specimens and thus this finding can be considered as a local crinoid Konservat Lagerstätte. Isocrinus (Chladocrinus) pehuenchensis new species is described from a single articulated specimen preserved in a silty calcareous concretion collected from a late Hauterivian concretion level of the Agua de la Mula Member. It is characterized by isotomous branching, eight or nine secundibrachials (IIBr), slender column ornamented with medial ridge of fine tubercles, interradius acuminated with fine tubercles on its tip, short noditaxis, and pentastellate columnals. It is associated with low-energy fall-out deposits in the offshore. The excellent state of preservation was due to an early cementation process by carbonate that enhanced lithification around the specimen.

UUID: http://zoobank.org/4763ae70-9d0a-4015-b1c9-6a7ceeedfb1a

Aerliceaster nexosus n. gen. n. sp. (Echinodermata), one of the oldest of known asteroids, is based on six specimens from the Floian (Early Ordovician) Garden City Formation of Idaho, and Kolataster perplexus n. gen. n. sp. is based on two specimens from the Sandian (Late Ordovician) Mifflin Formation of Illinois. Although the asterozoan skeleton is subdivided into few ossicular categories, evolutionary derivations of all the categories are not fully established, and therefore published evaluations differ. Beginning with phylogenetic work placing asteroid ancestry within the Somasteroidea together with the new taxa described herein, aspects of early asteroid morphology are evaluated and ambiguities in need of further study identified. Uncertainties are considered to be founded in rapid early asterozoan diversification and the scanty fossil record.

UUID: http://zoobank.org/b43d07cc-c8fb-4a84-bc6f-40aa6e0daea2

In contrast to almost all other invertebrate phyla that constructed biomineralized skeletons during the “Cambrian explosion” and maintained them during the entire fossil record, ascidian tunicates evolved this protective and stabilizing advantage only during the Permian, although soft-bodied representatives of this subphylum made their first appearance already in the early Cambrian. It remains enigmatic why these compound calcareous skeletons persisted only until the Late Triassic, subsequently followed by less-rigid internal skeletons from the Lower Jurassic onwards, which consist of scattered isolated spicules only. In addition to recently described aragonitic ascidian exoskeletons from the Permian and Triassic, new discoveries of similar, but colonial ascidian compound endoskeletons in the lower Carnian exhibit a short-living branch of this group, which moreover contain the first indubitable calcareous spicules. The latter are embedded in the solid endoskeleton, which is composed of polygonal aragonitic plates with smooth outer and zigzag lined inner boundaries. They consist of irregular, parallel (orthogonal), or fan-shaped (clinogonal) arrangements of acicular aragonite crystals. The following taxa are described as new: order Cassianomorpha new order with the family Cassianosomidae new family and the genus Toscanisoma new genus with the species T. multipartitum new species and T. triplicatum new species.

UUID: http://zoobank.org/03555353-cdab-42e8-8e99-9bfce15fa249

This report documents the last pteraspids, (armored, jawless members of the Heterostraci), which are otherwise only known from the Early Devonian of the Old Red Sandstone Continent. Tuberculate pteraspid heterostracans are described from the Middle Devonian beds of two formations in western North America. The late Givetian Yahatinda Formation of Alberta and British Columbia consists of channels cut into lower Paleozoic rocks and represents deposition in marine to littoral environments. Clavulaspis finis (Elliott et al., 2000a) new combination is redescribed from additional material from the Yahatinda Formation and reassigned to the new genus Clavulaspis because the original genus name is invalid. The Eifelian Spring Mountain beds of Idaho consist of a large channel that represents a clastic-dominated estuarine environment. It contains Scutellaspis wilsoni new genus new species, and the previously described species from the Spring Mountain beds is redescribed and reassigned to Ecphymaspis new genus, which was prompted by new material and a review of the validity of the original genus name. Phylogenetic analysis shows that these three new taxa form part of the derived clade Protaspididae.

UUID: http://zoobank.org/9cf09b21-cec1-4ce4-bc2b-658d0b515e10

Four placoderm fishes are described from the Lepel Beds of the Vitebsk Regional Stage of the Lower Devonian (upper Emsian) of Belarus. Two forms are new (Stipatosteus svidunovitchi new genus, new species and Actinolepis zaikai new species) and the other two taxa are left in open nomenclature (Coccosteidae gen. indet. sp. indet.; Placodermi indet. 1). Specimens of all four forms consist of disarticulated, mostly well-preserved plates (some are worn at the edges) originating from the cores of six boreholes. Additionally, the paper presents data on organic remains with the placoderms. A detailed lithological description of the enclosing rocks and a taphonomic description of the ichthyofauna are also provided. The 2010 Stratigraphic Chart of the Devonian deposits of Belarus was used as the stratigraphic basis. The presence of the genus Actinolepis Agassiz, 1844 in these deposits as well as in the Emsian of the Baltic region and Spitsbergen suggests a regional paleogeographic connection at this time.

UUID: http://zoobank.org/368b44df-e9e9-417c-a0e0-f7e138b7f4ec

Deinotheriidae Bonaparte, 1845 is a family of browsing proboscideans that were widespread in the Old World during the Neogene. From Miocene deposits in the Indian subcontinent, deinotheres are known largely from dental remains. Both large and small species have been described from the region. Previously, only small deinothere species have been identified from Kutch in western India. In the fossiliferous Tapar beds in Kutch, dental remains have been referred to the small species Deinotherium sindiense Lydekker, 1880, but the specimens are too fragmentary to be systematically diagnostic. Here, we describe a large p4 of a deinothere from the Tapar beds and demonstrate that it is morphologically most similar to Deinotherium indicum Falconer, 1845, a large species of deinothere, thereby confirming the identity of deinotheres at Tapar. Deinotherium indicum from Tapar is larger than other deinotheres identified from Kutch and is the first occurrence of the species in the region. This new specimen helps constrain the age of the Tapar beds to the Tortonian and increases the biogeographic range of this species—hitherto only known from two localities on the subcontinent. This specimen also highlights the morphological diversity of South Asian deinothere p4s and allows us to reassess dental apomorphies used to delimit Indian deinothere species. Lastly, we argue that by the late Miocene, small deinotheres in Kutch were replaced by the large Deinotherium indicum.

Conularina triangulata (Raymond, 1905), the genotype of Conularina Sinclair, 1942, is a rare, early Late Ordovician conulariid (Cnidaria, Scyphozoa; Van Iten et al., 2006) having three sides or faces instead of four (Sinclair, 1942, fig. 9; Van Iten, 1992, text-fig. 3E). Originally described from the Valcour Formation (early Sandbian; Dix et al., 2013) on Valcour Island, New York (Sinclair, 1942), C. triangulata has since been found in laterally equivalent strata of the upper Laval Formation (‘Upper Chazy’; Sinclair, 1942) in Laval, Québec, Canada (Sinclair, 1942). From this same unit and area, Sinclair (1942) erected three new, four-sided species of Conularina (C. irrasa, C. raymondi, and C. undosa), and he erected a single four-sided species (C. narrawayi) from the Ottawa Formation (now the Sandbian–Katian Ottawa Group; Dix et al., 2013) at Tétreauville (now Gatineau), Québec. Subsequently, Jerre (1994) reported the occurrence of two species of Conularina in the Upper Ordovician of Sweden. Jerre (1994) also proposed that Eoconularia? forensis Sinclair, 1946 from the Upper Ordovician Citadelle Formation (‘Quebec City’ Formation; Sinclair, 1946) in Québec City, Québec (Promontoire de Québec thrust sheet, Appalachian Humber Zone, Allochtonous Domain; Castonguay et al., 2002) is a species of Conularina.