The thallus of a new noncalcified dasycladalean alga, Wiartonella nodifera n. gen. n. sp., from the mid-Silurian Eramosa Lagerstätte of Ontario, Canada, comprises a narrow main axis with laterals in whorls (euspondyl). Laterals branch to the second order and show a distinct expansion (node) at the termination of first-order lateral segments. Morphologic differences between specimens are interpreted as ontogenetic stages similar to those displayed by extant Dasycladales, including late-stage shedding of higher-order lateral segments. Examination of reproductive functional morphology using biophysical modeling indicates that the expanded terminations of the first-order lateral segments probably are not homologous with gametophores, and that reproduction instead was either endospore or cladospore, with details of the ontogenetic sequence pointing to the latter. The distinctive lateral morphology displayed by this species adds to the list of morphological innovation achieved by dasycladalean algae during a significant burst of evolutionary activity that unfolded between the Middle Ordovician and late Silurian.

New samples of fusulinids collected in the Tengchong Block, western Yunnan, China, are systematically studied and presented here. The fusulinid fauna from the Xishancun section in the Shanmutang area is dominated by Chusenella and Nankinella, whereas that from the Shuangheyan area is composed mainly of Chusenella and Schwagerina. Both faunas are dated as Roadian–Capitanian (middle Permian). These new findings are integrated with fusulinid taxa reported earlier from the block to demonstrate the taxonomic features and paleogeographic significance of Permian fusulinids. The low generic diversity through early and middle Permian and the paucity of middle Permian neoschwagerinids and verbeekinids in the block confirm its Gondwana-affinity attributes. Moreover, the Permian fusulinids of the Tengchong Block are depauperate; i.e., consisting of a limited number of species with abundant individuals. This particular feature commonly suggests an inhospitable environment, and carbonates of varied facies containing these faunas in the Tengchong Block suggest a facies-independent factor as the reason, most likely the relatively low temperature of seawater.

Cherty limestones, marls and radiolarites are widespread in the Hallstatt Mélange of the Northern Calcareous Alps. The mélange was formed during the Neotethyan orogeny in a series of deep-water basins that progressively developed in front of the advancing nappe front. The low thermal overprint of these rocks favors the good preservation of radiolarians (polycystines) that have been used for dating and reconstructing the Jurassic tectonostratigraphy of the area. This paper describes rich Middle–Late Jurassic radiolarian faunas from four localities in the Hallstatt Mélange near Bad Mitterndorf in Austria. Two different successions, both spanning from the Bathonian to the Oxfordian, are dated. In the first succession, the radiolarites are intercalated between or occur as matrix in mass-flow deposits originating from the accretionary wedge. The second succession is nearly 100 m thick but is devoid of mass-flow deposits and documents a continuous radiolarite deposition in greater distance from the nappe stack. Both successions are ascribed to the Sandlingalm Basin, which evolved on a relatively distal continental margin during early stages of the orogeny.

The highly diverse and well-preserved radiolarian assemblages have been used for a detailed taxonomic study. Two new families are described: Minocapsidae n. fam. and Xitomitridae n. fam.; six new genera are described Doliocapsa n. gen., Crococapsa n. gen., Parvimitrella n. gen., Xitomitra n. gen., Campanomitra n. gen., and Mizukidella n. gen. In addition, one new replacement name, Takemuraella (pro Triversus), is introduced, the diagnoses of 6 genera are emended, and two new species are described: Hemicryptocapsa nonaginta n. sp. and Mizukidella mokaensis n. sp.

Although putative corals of uncertain affinities occur in the early Cambrian, the earliest definite tabulate corals have not been described prior to the Early Ordovician in North America. This paper reports a new finding of a tabulate-like coralomorph forming part of biostratigraphically well-constrained reef mounds in the latest Cambrian–Early Ordovician La Silla Formation in the Argentine Precordillera. The oldest record of the coralomorph genus Amsassia is reported and a new species, A. argentina, is erected. The discovery of this genus in the lowermost Ordovician modifies the previously proposed paleogeographic distribution and patterns of origination and migration routes of this coral-like organism. Amsassia argentina n. sp. constitutes a main framework builder together with a complex microbial consortium. This oldest occurrence of Amsassia as a reef builder represents a new record of a skeletal organism in the gap of metazoan reef constructors after the demise of archaeocyaths in the late early Cambrian.

New discoveries of the early Cambrian yiliangellinine trilobite Zhangshania typica Li and Zhang in Kunming preserve almost all instars from early postembryonic (protaspid) to mature (holaspid) phases in articulated state, in addition to mature specimens with antennae bearing paired spines on the basal articles. The ontogenetic series shows protarthrous development with some, but likely not all, early holaspid instars expressing additional pygidial segments, gradual rearward migration of the location of the longest pleural spines on the trunk segments, and striking positive allometry of the genal spines. It also reveals Parazhangshania sichuanensis Li and Zhang, 1990 to be the holaspid stage 3 of Z. typica, and therefore its junior synonym. This new find in the Hongjingshao Formation provides species-based regional correlation across the South China block and Z. typica may provide an important biostratigraphic marker for the base of the traditional Tsanglangpuan Stage.

Tiny, pelagic arthropods from the Anisian Luoping Biota exposed in two quarries near Luoping, Yunnan Province, China, represent the numerically most abundant organisms in the assemblage. They form the basis for definition of two, and possibly three, species referred to the order Lophogastrida, family Eucopiidae. Yunnanocopia grandis new genus new species and Y. longicauda n. gen. new species represent the oldest occurrence of mysidaceans in the fossil record. Their anatomy allies them with the Ladinian species Schimperella acanthocercus Taylor, Schram, and Shen, 2001, from Guizhou Province, China, which previously was thought to be the oldest lophogastrid, and with extant species of Eucopiidae. Their appearance in the Anisian represents one additional element of the early faunal radiation within the Luoping Biota following the end-Permian extinction event. Presence of well-preserved oostegites, along with other morphological features, documents a conservative bauplan expressed in Eucopiidae.

An exceptional fauna of retiolitine graptolites from Aeronian and lowermost Telychian strata in Arctic Canada provides significant new insights into the phylogeny and history of diversity of retiolitine graptolites. All specimens were isolated by dissolution of calcite concretions. The results indicate that retiolitines emerged within the lower Aeronian and reached a higher than expected level of diversity and disparity of forms by mid-Aeronian time. The uppermost Aeronian is almost totally devoid of preserved graptolites in Arctic Canada and, therefore, our material provides few new insights into retiolitine morphology or diversity through that interval. Specimens assigned to Pseudoretiolites? sp. occur in well-dated lower Aeronian strata, thus representing the lowest known biostratigraphic occurrences of retiolitines globally. This taxon appears to be morphologically primitive in that the sicula is completely preserved, with preservation of the proximal regions of theca 11, as well as distal thecal fusellum on mature specimens. Cladistic analysis of the Llandovery retiolitines shows that Pseudoretiolites is a stem genus for all of the other retiolitine taxa, which comprise two clades: one consisting of Pseudoplegmatograptus, Retiolites, and Stomatograptus and their derivatives; and the other includes Rotaretiolites, Aeroretiolites, Aeroretiolites?, Eorograptus, Paraplectograptus, Paraplectograptus?, and Sokolovograptus, and all of those other taxa that had previously been placed in the Plectograptinae. We follow the recent proposal that all of the taxa traditionally assigned to the Retiolitidae be assigned to the subfamily Retiolitinae. The following new taxa are described: Pseudoretiolites hyrichus n. sp., Eorograptus spirifer n. sp., and Aeroretiolites cancellatus n. gen. n. sp.

Ceratodontid lungfishes are generally rare, poorly represented elements of North America’s Mesozoic ecosystems, with previously known maximum diversity in the Late Jurassic. Herein we describe four new species of the form genus Ceratodus, from the Cretaceous of the Western Interior, considerably expanding fossil representation of post-Triassic dipnoans in North America. To model taxonomic and morphologic diversity, we adopt a four-fold system of phenetically based species groups, named for exemplars from the Morrison Formation. Ceratodus kirklandi n. sp. (Potamoceratodus guentheri group) and C. kempae n. sp. (C. frazieri group) represent a hitherto unsampled time interval, the Valanginian. Ceratodus nirumbee n. sp. and C. molossus n. sp. extend the temporal ranges of the C. fossanovum and C. robustus groups upward to the Albian and Cenomanian, respectively. These new occurrences show that ceratodontids maintained their highest diversity from the Late Jurassic through the mid-Cretaceous (Albian–Cenomanian), an interval of ~60 Myr. The existing record suggests that some of the later (mid-Cretaceous) ceratodontids may have been tolerant of salt water; to date, there is no evidence that they aestivated. Only a few occurrences are known from horizons younger than Cenomanian. Demise of ceratodontids appears to be part of a broader pattern of turnover that occurred at the Cenomanian-Turonian boundary in North America.

We describe a new species of the aetosaur Coahomasuchus, C. chathamensis, based on an incomplete, but largely articulated, anterior portion of a skeleton recovered from a quarry in the Upper Triassic Pekin Formation of Chatham County, North Carolina. This is only the second documented occurrence of Coahomasuchus, with the other being the holotype of C. kahleorum Heckert and Lucas, 1999 from the Upper Triassic Colorado City Formation of Texas. Although much of the specimen is the same size as the holotype of C. kahleorum, the dorsal paramedian osteoderms of the North Carolina taxon are considerably (~1.3×) wider than homologous counterparts in C. kahleorum, and the ventral thoracic osteoderms are also rectangular (~1.5× wider than long), rather than square, presumably to accommodate the wider body. This is a rare instance where two articulated specimens of closely related aetosaur species are available for direct comparison of homologous osteoderms. Isolated osteoderms with similar ornamentation from the same locality indicate that C. chathamensis may have been one of the earliest aetosaurs to attain the broad osteoderm proportions (width:length >3.5:1) otherwise known solely from later branching, spinose taxa such as Typothorax. The co-occurrence of Lucasuchus and Coahomasuchus in both North Carolina and Texas supports past correlations indicating an Otischalkian (Carnian) age for these strata and demonstrates that plesiomorphic, non-spinose aetosaur genera were not necessarily endemic to a single basin in North America.

Lower and upper third molars of a proboscidean from the Tha Chang sand pits, Nakhon Ratchasima, northeastern Thailand, show a zygodont pattern. The crescentoids are less well developed than those of the type specimen of Zygolophodon gobiensis but similar to those of late Miocene specimens from south China assigned to Z. lufengensis and Z. chinjiensis. On the other hand, the loph(id)s are less oblique and the zygodont crests are less well developed than in Z. lufengensis and Z. chinjiensis. However, it is difficult to erect a new species for these specimens because their anterior loph(id)s are so deeply worn that the morphology of the conelets on these loph(id)s is unclear. Thus, we identify the specimens as Zygolophodon sp. Zygolophodon is known from lower and middle Miocene sediments in Africa while they are usually dated to the early–late Miocene in Europe, South Asia such as Pakistan, and Central and North China. Based on its apparent grade of dental evolution, the new material of Zygolophodon from Thailand is probably late Miocene in age. In addition, this discovery is the first record of a zygodont proboscidean in Southeast Asia.

Waagen (1875) was the first who dealt with the Jurassic ammonites of Kutch based on detailed taxonomic work. In his monograph, he described among many species Perisphinctes spirorbis Neumayr, 1870 and P. aberrans Waagen, 1875 from the Callovian of Kutch, but the figures mentioned in the description did not correspond to the actual species. For P. spirorbis, the plate illustrated the holotype of an entirely different species (P. aberrans). On the other hand, P. spirorbis was illustrated as the lectotype of P. aberrans. Later, Spath (1924) introduced a new genus, Subgrossouvria, based on Waagen’s P. aberrans as type species. He also erected another genus, Indosphinctes, and included P. spirorbis Waagen, 1875 within the synonymy of I. indicus (Siemiradzki, 1899). Spath (1931) was aware of wrong numbering of plates of Waagen’s two species. But subsequent workers were ignorant of these taxonomic errors and continued to refer Waagen’s wrong plate numbers. We have here described both the type specimens and provided diagnoses for Spath’s two genera. We plead for this taxonomic correction in the incoming revised Treatise on Jurassic ammonites.