The effects of contrasting management systems either of infrequent
rotational grazing by town milk
supply dairy cattle, or of frequent defoliation by continuously
grazing sheep and beef cattle, on the
morphology of independent plants and populations of ‘Grasslands
Roa’ tall fescue in mixed
pastures, were measured over 1 year (1992/93) in New Zealand.
Volunteer perennial ryegrass plants were also measured for comparison.
While both species exhibited a similar pattern of clonal growth, tall
fescue developed more plants of
higher branching complexity than perennial ryegrass, chiefly through
maintaining more connective
stems, as herbage production was confined to the three youngest branching
orders in both species.
Greater resistance to microbial degradation of old stems through poorer
quality organic matter (wide
C[ratio ]N ratio) compared to perennial ryegrass may be responsible for
the greater complexity of fescue
plants. In addition, tillering rates in tall fescue were three times
lower which was offset by greater
longevity and size of leaves and tillers, compared to perennial ryegrass.
As a result, seasonal
fluctuation in the distribution of plants among the various branching
orders in tall fescue was small,
producing a more stable population relative to the distinct seasonal
changes in the population of
perennial ryegrass plants. Grazing management had no effect on the
seasonal population structure in either species.
Differences in plant structure due to grazing management were small,
with
only slightly more tillers
on sheep-grazed than on cattle-grazed tall fescue plants. The major effect
of grazing management was
on dry weight or size of plant components. Cattle-grazed tall fescue plants
were 120% heavier, with
greater numbers and lengths of stolons and flower heads than those
under sheep grazing. For the
volunteer perennial ryegrass, the difference was only 65%, possibly
due to competition from the more
vigorous tall fescue under rotational cattle grazing.
Both species produced stolons throughout the year, although these were
primarily associated with
reproductive growth in spring. In tall fescue, an additional distinction
was made between stolon and
rhizome, the latter occurring mainly in the summer–autumn.
Their possible functions in plant growth are discussed.