Non-marine and marine palynomorphs were collected from a Telychian through Gorstian rock outcrop located near Allenport, Pennsylvania, USA in the east-central part of the Appalachian Foreland Basin. Close sampling of the section revealed six distinct palynological assemblages of spores, cryptospores, acritarchs and prasinophycean algae, likely deposited in nearshore marine settings. The Silurian Period is a critical time in plant evolution as megafossils found elsewhere indicate that vascular plants (tracheophytes) originated during this interval. Palynoflorules generally show a transition from cryptospore-dominated to miospore-dominated assemblages during the latter half of the Silurian. This transition is evident at Allenport, where over 30 species of miospores and cryptospores are recognized, including two new species: Vermiverruspora cottera and Rugosphaera falloambita. The sequential progression of innovations in exine structure and sculpture seen in the nonmarine palynomorphs at Allenport closely matches evolutionary patterns documented previously in the Appalachian Basin, Avalonia, and elsewhere around the world.

†Notogoneus osculus Cope, the type species for the geographically widespread Upper Cretaceous to Upper Oligocene genus †Notogoneus (Gonorynchidae), is redescribed in detail based on much new material. This species is known from the Early Eocene Fossil Butte Member of the Green River Formation, southwestern Wyoming. It is geologically the youngest known species of the family Gonorynchidae in North America, although the family is known from Oligocene and Miocene lacustrine deposits of Europe and Australia and is extant today in tropical marine environments of the Pacific (Gonorynchus spp.).

In this paper we also correct a number of mistakes in previously published morphological descriptions of †N. osculus (e.g., in skull and caudal skeleton) based on a large, new sample of well-preserved, well-prepared specimens. Accurate description of this species is important because it is the type for the genus and it is also by far the best preserved of all known species of †Notogoneus. In addition, it may shed new light on the interrelationships of Gonorynchidae (work in progress by the authors).

†Notogoneus appears to have inhabited a freshwater, subtropical environment. Based on distribution data and species diversity within the Green River Formation, we suspect that †N. osculus may have been a migratory fish, living in the central (off-shore) regions of Fossil Lake during part of its life, and in connecting streams and rivers during other parts of its life. Most or all species of the genus †Notogoneus are known from freshwater deposits and species are known from North America, Europe, Asia and Australia. Species of the extant gonorynchid genus Gonorynchus are all marine and are known from the Indo-Pacific and southern Atlantic.

Specimens of the large, shallow-marine, volutid gastropod Volutoderma Gabb, 1877, herein recognized only from strata of Late Cretaceous (Coniacian through early Maastrichtian) age in British Columbia, Washington, California, and Baja California have commonly been identified as Volutoderma averillii (Gabb, 1864). This review of available specimens assigns them to two genera: Volutoderma and Longoconcha Stephenson, 1941.

Twelve species, nine of them new, comprise three morphologic lineages of Volutoderma, i.e., 1) “Typical” includes V. querna n. sp., V. averillii (Gabb), V. blakei n. sp., V. jalama n. sp., V. perissa n. sp., and possibly Volutoderma? n. sp.; 2) “Angelica” includes V. angelica n. sp., V. elderi n. sp., and V. ynezae n. sp.; and 3) “Magna” includes V. santana Packard, V. magna Packard, and perhaps V.? antherena n. sp. A new species of Longoconcha, L. eumeka, is the first Pacific Slope record of this genus, which has a Gulf Coast and Tethyan Old World distribution. A smaller volutid, Retipirula Dall, 1907 is endemic to the study area and was formerly known only from its type species R. crassitesta (Gabb, 1869) of Paleocene age. Two new Retipirula are reported: R. calidula of latest Maastrichtian age and R. pinguis of Paleocene age.

Only the Volutoderma lineage containing V. averillii has been found north of San Francisco. Recovery of rudist bivalves from formations yielding Volutoderma suggests that these volutes were warm-temperate to subtropical gastropods. Co-occurrences of these gastropods and rudistids may aid in placing the warm-temperate/subtropical boundary during the Late Cretaceous.

Two new Deseadan interatheriine genera (Interatheriidae, Notoungulata) from the late Oligocene Salla Beds of BOLIVIA are described. Both are monotypic and one is known from a partial skeleton, a rarity among known pre-Santacrucian interathere taxa. Phylogenetically, both taxa nest well within Interatheriinae, showing characteristically bilobed p3-4. Both taxa also have derived characters (hypselodont cheeckteeth, persistent lingual sulcus on upper molars) relative to basal interatheriines such as Santiagorothia and Proargyrohyrax but are clearly plesiomorphic with respect to younger, more highly derived Santacrucian interatheriine taxa such as Interatherium and Protypotherium. New species Brucemacfaddenia boliviensis is on average larger than the other new Salla interatheriine, Federicoanaya sallaensis, although they do overlap in size. Distinguishing between the two new taxa based purely on molar morphology is confounded by lack of diagnostic characters on the molar teeth and the overlap in size between the taxa. We overcome this difficulty of identifying specimens that preserve only molars by using discriminant analysis. We present a few of the simpler yet still robust discriminant functions we used so that future workers have a means of identifying problematic specimens. Analysis of Salla interathere specimens and stratigraphic provenance indicates both taxa experienced a modest increase in body size upsection, the driving mechanism for which remains unknown, but could be environmental changes or simple drift. These two new taxa help emphasize the fact that while the Salla fauna shares elements with roughly contemporaneous Deseadan faunas from more southerly latitudes, important faunal distinctions mark the two regions as well.

A new genus and species of Asteroidea (Echinodermata), Embolaster graffhami, Embolasteridae, n. fam., is described from the Upper Ordovician Bromide Formation of Oklahoma. Embolaster is important in part because of the presence of two similar ossicles on each interbrachial plane in the axillary position; only a single axillary is recognized in most Paleozoic asteroids. the axillary traditionally has been considered to be a part of the marginal series, although marginals are almost always paired. the two ossicles are interpreted as axillaries and the homologues of an ancestral proximal marginal pair. the odontophore, which lies distal to the mouth angle pair of most asteroids, tentatively is the homologue of the proximal axillary.

Embolaster also is important because of the presence of an ossicular series between the ambulacrals and the putative adambulacrals. An extra ossicular series in this position is known from taxa traditionally assigned to the ophiuroids; however, other characters of Embolaster indicate asteroid affinities for this genus. the two ossicles lateral to and aligned with the ambulacrals in Embolaster are interpreted as homologous with the similarly arranged radial virgal series of somasteroids, although specific ossicular homologies are problematic.

A part of the mouth frame structure of Embolaster is exposed, although it is tiny and therefore difficult to interpret. the mouth frame arrangement is reminiscent of those of somasteroids, with small, narrow circumorals and apparent buccal podial basins directed toward the mouth opening. Many of the striking characters of Embolaster probably are plesiomorphic; apomorphies are difficult to ascertain.

Newly found fossils in the Conception and St. John's groups of the Bonavista Peninsula considerably extend the known geographic distribution of the Ediacaran fossils in Newfoundland. They occur in deepwater sediments and are preserved as epireliefs, forming census populations underneath volcanic ash layers throughout a more than 1 km thick turbiditic sequence. the exposed fossiliferous units comprise the Mistaken Point, Trepassey, Fermeuse, and Renews Head formations. the remains are tectonically deformed, with long axes of elliptical discs aligned parallel to cleavage strike; shortening of originally circular bedding surface features is on the order of 30-50% (averaging ~35%).

The assemblage includes Aspidella, Blackbrookia, Bradgatia, Charnia, Charniodiscus, Fractofusus, Hiemalora, and Ivesheadia. These occur throughout the succession, with Aspidella being the most common genus, followed by Charnia and Charniodiscus. Four new taxa are described, with candelabra-like fossils with a Hiemalora-like base referred to Primocandelabrum hiemaloranum n. gen. and sp., bush-like fossils to Parviscopa bonavistensis n. gen. and sp., ladder-like fossils to Hadryniscala avalonica n. gen. and sp., and string-like fossils with basal disc to Hadrynichorde catalinensis n. gen. and sp. the remains also include dubiofossils. The stratigraphic ranges of some taxa on the Bonavista Peninsula are longer than previously reported from the Avalon Peninsula, with Fractofusus spindles present in the Trepassey Formation, Bradgatia, Charnia, Charniodiscus, and Ivesheadia reaching as high as the Fermeuse Formation, and Aspidella extending into the middle of the Renews Head Formation. the spindles in the Trepassey Formation are comparable to those found mainly in the stratigraphically older Briscal Formation on the Avalon Peninsula.

Brachiopods of the orders Strophomenida, Orthotetida, and Athyridida from the late middle Frasnian–early Famennian interval (hassi to triangularis conodont zones) in the Namur-Dinant Basin (southeastern margin of Laurussia; southern Belgium) are described. Nine genera and subgenera represented by 12 species are recognized here. One new genus, Retrorstrophia, and one new species, Douvillina area, are erected. Crinisarina stainbrooki is proposed to solve the homonymy between C. reticulata (Gosselet, 1877) and Cleiothyridina reticulata Stainbrook, 1947, the type species of Crinisarina Cooper and Dutro, 1982. A lectotype is selected and illustrated for C. reticulata. A new name is also proposed for Athyris reticulata Chen and Xu, 2000, which is assigned here to Crinisarina: C. shashishanensis nom. nov. In southern Belgium, the last representatives of the families Douvillinidae and Leptostrophiidae (Strophomenida) are from the Upper rhenana Zone (late Frasnian). Athyridid brachiopods, especially the representatives of the subfamily Cleiothyridininae constituted a significant part of the early Famennian recovery faunas with spiriferids and rhynchonellids, just after the late Frasnian mass extinction.

The Middle Cambrian Spence Shale Member (Langston Formation) and Wheeler and Marjum Formations of Utah are known to contain a diverse soft-bodied fauna, but important new paleontological material continues to be uncovered from these strata. New specimens of anomalocaridids include the largest and smallest near complete examples yet reported from Utah. New material of stem group arthropods includes two new genera and species of arachnomorphs: Nettapezoura basilika and Dicranocaris guntherorum. Other new arachnomorph material includes a new species of Leanchoilia comparable to L. protogonia Simonetta, 1970; Leanchoilia superlata? Walcott, 1912; Sidneyia Walcott, 1911a; and Mollisonia symmetrica Walcott, 1912. L. protogonia from the Burgess Shale is confirmed as a separate species and is not a composite fossil. The first example of the trilobite Elrathia kingii preserving traces of the appendages is described. In addition, new material of the bivalved arthropods Canadaspis Novozhilov in Orlov, 1960; Branchiocaris Briggs, 1976; Waptia Walcott, 1912; and Isoxys Walcott, 1890 is described.

The Cambrian inlier at Beaver Harbour, southern New Brunswick, is now confidently referred to the marginal platform of the late Proterozoic–Early Paleozoic Avalon microcontinent. The sub-trilobitic Lower Cambrian Chapel Island and Random Formations are unconformably overlain by the mafic volcanic-dominated Wade's Lane Formation (new). Late Early Cambrian trilobites and small shelly taxa in the lowest Wade's Lane demonstrate a long Random–Wade's Lane hiatus (middle Terreneuvian–early Branchian). Latest Early–middle Middle Cambrian pyroclastic volcanism produced a volcanic edifice at Beaver Harbour that is one of three known volcanic centers that extended 550 km along the northwest margin of Avalon. Middle Middle Cambrian sea-level rise, probably in the Paradoxides eteminicus Chron, mantled the extinct volcanics with gray-green mudstone and limestone of the Fossil Brook Member. Black, dysoxic mudstone of the upper Manuels River Formation (upper Middle Cambrian, P. davidis Zone) is the youngest Cambrian unit in the Beaver Harbour inlier. Lapworthella cornu (Wiman, 1903) emend., a senior synonym of the genotype L. nigra (Cobbold, 1921), Hyolithellus sinuosus Cobbold, 1921, and probably Acrothyra sera Matthew, 1902a, range through the ca. 8 m.y. of the trilobite-bearing upper Lower Cambrian, and H. sinuosus and A. sera persist into the middle Middle Cambrian. Lapworthella cornu and H. sinuosus replaced the tropical taxa L. schodackensis (Lochman, 1956) and H. micans Billings, 1872, in cool-water Avalon.

Radiolarians collected from five localities of the Baltalimani Formation in the Istanbul region are described in detail, and the local stratigraphic ranges and persistence of biostratigraphically important radiolarian species are documented. the spumellarian component is published for the first time, and four new species are described: Albaillella kayai n. sp., Ceratoikiscum jucundum n. sp., Meschedea akcetensis n. sp., and Stigmosphaerostylus istanbulensis n. sp. Two assemblages are recognized: the A. paradoxa and the A. sp. aff. A. undulata-A. deflandrei assemblage. the older A. paradoxa assemblage contains Traenosphaera sicarius, Pararchocyrtium serrensis, Cyrtisphaeractenium mendax, and common to abundant A. paradoxa and Popofskyellium. Albaillella paradoxa is not restricted to the older assemblage but is the only species of Albaillella present. the younger A. sp. aff. A. undulata-A. deflandrei assemblage contains either A. sp. aff. A. undulata or A. deflandrei. Albaillella indensis ambigua, A. kayai n. sp., and Kantollium sp. A occur exclusively in the A. sp. aff. A. undulata-A. deflandrei assemblage, but are rare and patchy in their occurrence. Distinctive taxa common to both assemblages are Archocyrtium lagabriellei, Pylentonema antiqua, C. avimexpectens, C. umbraculatum, C. jucundum n. sp., Trilonche palimbola, M. akcetensis n. sp., S. tortispina, Polyentactinia polygonia, P. aranea, and Cubaxonium? octaedrospongiosum. Many of the taxa present occur in middle and upper Tournasian sections from France, the central Pyrenees, and Germany and provide a solid basis for re-assessing the age of the Baltalimani Formation as middle to late Tournasian (Tn2-Tn3).

Acid processing allowed systematic identification of 458 Upper Triassic silicified scleractinian corals (20 genera, 47 species) from the Alexander terrane (southeast Alaska) and Wrangellia (Wrangell Mountains, southern Alaska and Vancouver Island, Canada). Coral faunas, here presented, show taxonomic affinity with coeval collections from other Cordilleran terranes, specifically the Wallowa terrane (northeastern Oregon and Idaho) and Peru (South America) as well as the distant Tethys region. Genera from the Alexander terrane include: Kompsasteria Roniewicz, Gablonzeria Cuif, Cuifia Melnikova, Paracuifia Melnikova, Distichophyllia Cuif, Retiophyllia Cuif, Kuhnastraea Cuif, Margarosmilia Volz, Distichomeandra Cuif, Astraeomorpha Reuss, Pamiroseris Melnikova, Crassistella Roniewicz, Stylophyllum Frech, and Meandrostylis Frech. Genera from Wrangellia include: Gablonzeria Cuif, Distichophyllia Cuif, Retiophyllia Cuif, Kuhnastraea Cuif, Margarosmilia Volz, Distichomeandra Cuif, Astraeomorpha Reuss, Parastraeomorpha Roniewicz, Chondrocoenia Roniewicz, Pamiroseris Melnikova, Crassistella Roniewicz, Ampakabastraea? Alloiteau, Recticostastraea Stanley and Whalen, Meandrostylis Frech, Anthostylis Roniewicz, and the new genus Campesteria n. gen. New species include: Gablonzeria grandiosa n. sp., Paracuifia smithi n. sp., Paracuifia jennieae n. sp., P. anomala n. sp., Retiophyllia dendriformis n. sp., R. obtusa n. sp., and Campesteria prolixia n. sp.

Systematics of the trilobite subfamily Koneprusiinae Vaněk and Pek, 1987 are revised on the basis of new material and a comprehensive parsimony analysis. New taxa include the genus Nevadaprusia (type species N. cortezensis n. sp.) from the Pragian of Nevada and the species Laethoprusia graffhami n. sp. from the Lochkovian of Oklahoma, L. cozarti n. sp. from the Lochkovian of Tennessee, and Laethoprusia n. sp. A from the Sheinwoodian of Arctic Canada and the southern Mackenzie Mountains. Nevadaprusia insolita (Haas) is revised on the basis of new material. Laethoprusia and Nevadaprusia are interpreted as sister taxa, and Koneprusia and Isoprusia are successive sister groups to the Laethoprusia/Nevadaprusia clade.

The biostratigraphy of the upper part of the Mississippian Lower Limestone Formation in the Midland Valley, Scotland is revised using foraminiferal assemblages. This formation was previously assigned exclusively to the upper part of the Brigantian Substage (upper Cf6δ, P2 or latest Visean), whereas the succeeding Limestone Coal Formation (barren of foraminifers and conodonts) was assigned to the Pendleian Substage (lower Cf7, E1 or early Serpukhovian). The foraminifers, in particular those recorded from the Second Hosie and Top Hosie limestones and their lateral equivalents (Anvil and MacDonald limestones), are comparable to Serpukhovian assemblages from the Ukraine and Russia. Consequently, the Visean/Serpukhovian stage boundary is repositioned at the base of the Second Hosie Limestone. This assignment of the Second Hosie Limestone to the Pendleian is reinforced by the first occurrence of the ammonoid Emstites (Cravenoceras), although it occurs 1 m below the Top Hosie Limestone. Pendleian foraminiferal assemblages from northern England, its geographically closest equivalent, are similar but not well enough known for a detailed comparison. A few similarities are found between the foraminiferal assemblages from the Midland Valley and those from southwest Spain and North African basins. Faunas from the Second and Top Hosie limestones and their lateral equivalents allow us to propose an assemblage zone, valid for regional correlations within the British Isles, and the faunas can be potentially used as Serpukhovian markers for western Paleotethyan basins. Two new genera and species are described, Praeplectostaffella anvilensis n. gen. n. sp. and Praeostaffellina macdonaldensis n. gen. n. sp., and two new species, Tubispirodiscus hosiensis n. sp. and Euxinita pendleiensis n. sp.

A variety of Pennsylvanian sponges have been recovered from exposures along the shore of Lake Bridgeport and in nearby areas in Wise County, north-central Texas. Calcareous and hexactinellid sponges have locally weathered out of the Jasper Creek Shale (= upper Lake Bridgeport Shale) and the overlying Devil's Den Limestone of the Graford Formation, of Late Pennsylvanian Missourian age.

The Demospongea protomonaxonids Heliospongia excavata King, 1933, and Coelocladia spinosa Girty, 1908 are represented in the studied collections by several specimens, as is the new genus and species Luterospongia texana. Agelasid ceractinomorph demosponges are represented by the fissispongiid Fissispongia jacksboroensis King, 1938, the maeandrostiid Maeandrostia kansasensis Girty, 1908, and the girtycoeliid, Girtycoelia typica King, 1933. Sponges of the Class Hexactinellida and the amphidiscophorid Family Stiodermatidae are represented by an extensive suite of specimens of the new genus and species Dermosphaeroidalis irregularis. Representatives of the hexactinellid reticulosid sponges include some unusually large specimens of the vase-shaped Endoplegma calathus Finks, 1960, of the Family Docodermatidae, and fragmental specimens of ?Stereodictyum orthoplectum Finks, 1960, of the Family Stereodictyidae. One unclassified root tuft fragment is also described and illustrated.

Much of this diverse sponge fauna was endemic to the western embayment of Pangaea.

locrinus trentonensis Walcott, 1883 is characterized by the widest food grooves and the largest covering plates of any of the Walcott-Rust Quarry crinoids, which indicates that the animal captured relatively large food particles with large and widely separated tubefeet. Although iocrinids are generally considered as primitive disparids, their anal sac is unique. the holdfasts of I. trentonensis consist of distal stem coils that are tightly wrapped around the columns of other crinoids. the relatively long column of Ectenocrinus simplex (Hall, 1847) was attached to a wide range of shelly substrates by a small irregular and somewhat lobate holdfast. Ectenocrinids ate much smaller food items that were collected by smaller and more tightly packed tubefeet. the ontogeny of E. simplex illustrates the differences between the food gathering systems of conspecific crinoids from shallow and deep water habitats. the calceocrinid Calceocrinus barrandii Walcott, 1883 lived with its long stem forming a runner along the seafloor. the crown was movably hinged to the basal circlet and the stem. Moderately wide food grooves were probably present.

Twenty-three species of Campanian, Maastrichtian, Paleocene, and Eocene freshwater gastropods from northeastern Mexico are described, including eight new species: Viviparus mcbridei, Pachychilus (Pachychiloides) lawtoni, Melanoides (Melanoides) yolandae, Melanoides (Melanoides) wollebeni, Physa cepedaensis, Mesolanistes magnus, Mesolanistes murrayi, and Gyraulus zoltani. Specimens were collected from fine-grained, green sandstone and mudstone, red mudstone, and as hematized remains in fine light-brown sandstones belonging to the following stratigraphic units of northeastern Mexico: Cerro del Pueblo (Campanian, Parras Basin), Olmos (Maastrichtian, Sabinas Basin), Las Encinas (Paleocene, Parras Basin), and Carroza (Eocene, La Popa Basin) Formations. All except two of the genera (?Pyrgulifera and Mesolanistes) have recent representatives. Review of habitats of living species of the six extant genera and interpretations of sample lithologies support the inference of dominantly freshwater paleoenvironments. Only one Maastrichtian locality is interpreted confidently to have had a brackish water influence.

We present a critical review of the alpha taxonomy and evolution of Eocene North American paromomyid primates, based on analysis of more than 570 stratigraphically controlled dental and gnathic specimens from the early Eocene of the southern Bighorn Basin, Wyoming (Wasatchian, Willwood Formation). In addition to documenting numerous previously unpublished specimens of known taxa (including deciduous teeth), we also describe a new species, Phenacolemur willwoodensis n. sp., from the upper part of the Willwood Formation (Wa 5 and 6). The new species is intermediate in size between Phenacolemur simonsi and Phenacolemur citatus and has both primitive features (e.g., retention of m2–3 paraconids, relatively long molar trigonids) and derived traits (e.g., relatively reduced paraconid on m1 and no p4 paracristid, unlike Paromomys). Overall patterns of dental evolution in southern Bighorn Basin paromomyids provide some support for previously hypothesized periods of faunal change (Biohorizons). In particular, Phenacolemur praecox evolves into the similarly sized but morphologically distinct Phenacolemur fortior at Biohorizon A, and P. fortior is replaced by P. citatus just below Biohorizon B. Two taxa previously believed to have become extinct at Biohorizon A (Ignacius graybullianus, P. simonsi) are shown to have persisted about a million years longer than previously thought. The Bighorn Basin paromomyids are of general interest in comprising a very dense sample that allows for the study of patterns of evolution against the backdrop of well-understood patterns of change in other mammalian lineages, and in climatic variables.

From the Aseelah Unit (Saal Formation), Batain Coast (eastern Oman), 48 bryozoan species were described. Eight species of them are new: cystoporids Cystodictya augusta n. sp. and Sulcoretepora orientalis n. sp., trepostomid Ulrichotrypella omanica n. sp., rhabdomesids Streblotrypa (Streblotrypa) binodata n. sp., Rhabdomeson coniforme n. sp., R. latum n. sp., Orthopora punctata n. sp., and phyloporine Rhombocladia quasiminor n. sp. A new combination is erected: Streblotrypa (Strebloscopora) katoi Sakagami, 2000 n. comb. Fifteen species are described in open nomenclature. the fauna shows mainly Lower to Middle Permian age, and paleobiogeographic connections to the Urals, southeast Asia and Australia.